SEC FORM 13F-HR

The Securities and Exchange Commission has not necessarily reviewed the information in this filing and has not determined if it is accurate and complete.
The reader should not assume that the information is accurate and complete.

UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549
FORM 13F

FORM 13F COVER PAGE

OMB APPROVAL
OMB Number: 3235-0006
Expires: July 31, 2015
Estimated average burden
hours per response: 23.8

Report for the Calendar Year or Quarter Ended: 09-30-2019
Check here if Amendment    Amendment Number:
This Amendment (Check only one.):    is a restatement.
   adds new holdings entries.
Institutional Investment Manager Filing this Report:
Name: US BANCORP \DE\
Address: U.S. BANCORP
800 NICOLLET MALL
MINNEAPOLIS, MN  55402-7020
Form 13F File Number: 028-00551

The institutional investment manager filing this report and the person by whom it is signed hereby represent that the person signing the report is authorized to submit it, that all information contained herein is true, correct and complete, and that it is understood that all required items, statements, schedules, lists, and tables, are considered integral parts of this form.

Person Signing this Report on Behalf of Reporting Manager:
Name: Michele Michals
Title: Vice President
Phone: 314-418-1707
Signature, Place, and Date of Signing:
Michele Michals Saint LouisMO 11-08-2019
[Signature] [City, State] [Date]
Report Type (Check only one.):
X 13F HOLDINGS REPORT. (Check here if all holdings of this reporting manager are reported in this report.)
   13F NOTICE. (Check here if no holdings reported are in this report, and all holdings are reported by other reporting manager(s).)
   13F COMBINATION REPORT. (Check here if a portion of the holdings for this reporting manager are reported in this report and a portion are reported by other reporting manager(s).)
Form 13F Summary Page
Report Summary:
Number of Other Included Managers: 2
Form 13F Information Table Entry Total: 6,951
Form 13F Information Table Value Total: 36,694,789
(thousands)
List of Other Included Managers:
Provide a numbered list of the name(s) and Form 13F file number(s) of all institutional investment managers with respect to which this report is filed, other than the manager filing this report.
[If there are no entries in this list, state “NONE” and omit the column headings and list entries.]
No. Form 13F File Number Name
1 028-02267 US BANK NATIONAL ASSOCIATION
5 028-16147 U.S. BANCORP INVESTMENTS, INC.

SEC FORM 13-F Information Table

The Securities and Exchange Commission has not necessarily reviewed the information in this filing and has not determined if it is accurate and complete.
The reader should not assume that the information is accurate and complete.

UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549
FORM 13F

FORM 13F INFORMATION TABLE

OMB APPROVAL
OMB Number: 3235-0006
Expires: July 31, 2015
Estimated average burden
hours per response: 23.8

COLUMN 1 COLUMN 2 COLUMN 3 COLUMN 4 COLUMN 5 COLUMN 6 COLUMN 7 COLUMN 8
VALUE SHRS OR SH/ PUT/ INVESTMENT OTHER VOTING AUTHORITY
NAME OF ISSUER TITLE OF CLASS CUSIP (x$1000) PRN AMT PRN CALL DISCRETION MANAGER SOLE SHARED NONE
DEUTSCHE BANK AG COMMON STOCK D18190898 159 21,265 SH   DFND 1 17,709 0 3,556
DEUTSCHE BANK AG COMMON STOCK D18190898 44 5,959 SH   OTR 1 3,413 1,978 568
ADIENT PLC COMMON STOCK G0084W101 89 3,855 SH   DFND 1 3,531 0 324
ADIENT PLC COMMON STOCK G0084W101 11 480 SH   OTR 1 457 23 0
ALKERMES PLC COMMON STOCK G01767105 6 287 SH   DFND 1 122 0 165
ALLEGION PLC COMMON STOCK G0176J109 1,385 13,360 SH   DFND 1 13,331 0 29
ALLEGION PLC COMMON STOCK G0176J109 560 5,403 SH   OTR 1 4,157 1,233 13
ALLERGAN PLC COMMON STOCK G0177J108 5,153 30,615 SH   DFND 1 17,365 0 13,250
ALLERGAN PLC COMMON STOCK G0177J108 383 2,276 SH   OTR 1 2,036 28 212
ALLERGAN PLC COMMON STOCK G0177J108 8 48 SH   DFND 5 48 0 0
AMCOR PLC COMMON STOCK G0250X107 8,202 841,229 SH   DFND 1 839,111 0 2,118
AMCOR PLC COMMON STOCK G0250X107 4,269 437,833 SH   OTR 1 395,719 192 41,922
AMCOR PLC COMMON STOCK G0250X107 87 8,990 SH   DFND 5 8,990 0 0
AMDOCS LTD COMMON STOCK G02602103 2,143 32,409 SH   DFND 1 32,332 0 77
AMDOCS LTD COMMON STOCK G02602103 246 3,723 SH   OTR 1 2,575 108 1,040
AMBARELLA INC COMMON STOCK G037AX101 900 14,321 SH   DFND 1 14,321 0 0
AMBARELLA INC COMMON STOCK G037AX101 92 1,463 SH   OTR 1 1,463 0 0
AMBARELLA INC COMMON STOCK G037AX101 12 206 SH   DFND 5 206 0 0
AON PLC COMMON STOCK G0408V102 2,619 13,524 SH   DFND 1 12,645 0 879
AON PLC COMMON STOCK G0408V102 3,255 16,813 SH   OTR 1 6,804 870 9,139
AON PLC COMMON STOCK G0408V102 28 145 SH   DFND 5 145 0 0
ARCH CAP GROUP LTD COMMON STOCK G0450A105 1,112 26,509 SH   DFND 1 18,429 0 8,080
ARCH CAP GROUP LTD COMMON STOCK G0450A105 561 13,372 SH   OTR 1 11,268 0 2,104
ARCH CAPITAL GROUP LTD COMMON STOCK G0450A105 36 878 SH   DFND 5 878 0 0
ARCOS DORADOS HOLDINGS INC A COMMON STOCK G0457F107 7 1,000 SH   OTR 1 1,000 0 0
ARGO GROUP INTL HLDGS LTD COMMON STOCK G0464B107 2,600 37,013 SH   DFND 1 19,413 0 17,600
ARGO GROUP INTL HLDGS LTD COMMON STOCK G0464B107 326 4,643 SH   OTR 1 3,912 731 0
ARGO GROUP INTERNATIONAL COMMON STOCK G0464B107 13 198 SH   DFND 5 198 0 0
ASSURED GUARANTY LTD COMMON STOCK G0585R106 278 6,258 SH   DFND 1 5,732 0 526
ASSURED GUARANTY LTD COMMON STOCK G0585R106 317 7,127 SH   OTR 1 0 30 7,097
ASSURED GUARANTY LTD COMMON STOCK G0585R106 2 46 SH   DFND 5 46 0 0
ATLASSIAN CORP PLC CLASS A COMMON STOCK G06242104 59 473 SH   DFND 1 325 0 148
ATLASSIAN CORP PLC CLASS A COMMON STOCK G06242104 72 578 SH   OTR 1 254 298 26
ATHENE HOLDING LTD CLASS A COMMON STOCK G0684D107 84 2,006 SH   DFND 1 2,006 0 0
ATHENE HOLDING LTD CLASS A COMMON STOCK G0684D107 122 2,881 SH   OTR 1 0 91 2,790
AXIS CAPITAL HOLDINGS LTD COMMON STOCK G0692U109 20 301 SH   DFND 1 231 0 70
AXIS CAPITAL HOLDINGS LTD COMMON STOCK G0692U109 5 77 SH   OTR 1 77 0 0
AXALTA COATING SYSTEMS LTD COMMON STOCK G0750C108 20 663 SH   DFND 1 450 0 213
AXALTA COATING SYSTEMS LTD COMMON STOCK G0750C108 2 51 SH   OTR 1 0 0 51
AXALTA COATING SYSTEMS LTD COMMON STOCK G0750C108 40 1,330 SH   DFND 5 1,330 0 0
ATLANTICA YIELD PLC COMMON STOCK G0751N103 1,868 77,576 SH   DFND 1 77,484 0 92
ATLANTICA YIELD PLC COMMON STOCK G0751N103 446 18,515 SH   OTR 1 18,415 0 100
ATLANTICA YIELD PLC COMMON STOCK G0751N103 8 359 SH   DFND 5 359 0 0
BANK OF NT BUTTERFIELD SON COMMON STOCK G0772R208 109 3,666 SH   DFND 1 0 0 3,666
BIOHAVEN PHARMACEUTICAL HOLD COMMON STOCK G11196105 0 10 SH   DFND 1 10 0 0
ACCENTURE PLC CL A COMMON STOCK G1151C101 188,278 978,830 SH   DFND 1 939,920 0 38,910
ACCENTURE PLC CL A COMMON STOCK G1151C101 57,531 299,094 SH   OTR 1 272,789 13,955 12,350
ACCENTURE PLC COMMON STOCK G1151C101 624 3,246 SH   DFND 5 3,246 0 0
BLUE CAPITAL REINSURANCE HOL COMMON STOCK G1190F107 11 1,500 SH   DFND 1 1,500 0 0
MIMECAST LTD COMMON STOCK G14838109 591 16,580 SH   DFND 1 16,490 0 90
MIMECAST LTD COMMON STOCK G14838109 185 5,200 SH   OTR 1 5,200 0 0
BROOKFIELD BUSINESS PARTNERS L COMMON STOCK G16234109 3 72 SH   DFND 1 72 0 0
BROOKFIELD PROPERTY PARTNERS L COMMON STOCK G16249107 2 103 SH   DFND 1 103 0 0
BROOKFIELD PROPERTY PARTNERS L COMMON STOCK G16249107 140 6,920 SH   OTR 1 6,920 0 0
BROOKFIELD INFRASTRUCTURE PART COMMON STOCK G16252101 75 1,520 SH   DFND 1 1,520 0 0
BROOKFIELD INFRASTRUCTURE PART COMMON STOCK G16252101 122 2,462 SH   OTR 1 1,000 0 1,462
BROOKFIELD RENEWABLE PARTNERS COMMON STOCK G16258108 41 1,000 SH   DFND 1 0 0 1,000
BROOKFIELD RENEWABLE PARTNERS COMMON STOCK G16258108 81 2,000 SH   OTR 1 2,000 0 0
BUNGE LIMITED COMMON STOCK G16962105 357 6,300 SH   DFND 1 6,300 0 0
CAPRI HOLDINGS LTD COMMON STOCK G1890L107 8 240 SH   DFND 1 71 0 169
CAPRI HOLDINGS LTD COMMON STOCK G1890L107 3 101 SH   OTR 1 101 0 0
CARDTRONICS PLC COMMON STOCK G1991C105 15 483 SH   DFND 1 483 0 0
CARDTRONICS PLC COMMON STOCK G1991C105 39 1,305 SH   OTR 1 0 1,305 0
CHINA YUCHAI INTL LTD COMMON STOCK G21082105 13 975 SH   DFND 1 756 0 219
CHINA BIOLOGIC PRODUCTS HOLD COMMON STOCK G21515104 1 13 SH   OTR 1 13 0 0
CREDICORP LTD COMMON STOCK G2519Y108 258 1,237 SH   DFND 1 1,038 0 199
CREDICORP LTD COMMON STOCK G2519Y108 27 128 SH   OTR 1 68 0 60
COCA COLA EUROREAN PARTNERS COMMON STOCK G25839104 10,670 192,423 SH   DFND 1 190,640 0 1,783
COCA COLA EUROREAN PARTNERS COMMON STOCK G25839104 2,477 44,676 SH   OTR 1 43,280 859 537
COCA-COLA EUROPEAN PARTNERS COMMON STOCK G25839104 59 1,074 SH   DFND 5 1,074 0 0
DELPHI TECHNOLOGIES PLC COMMON STOCK G2709G107 6 450 SH   DFND 1 450 0 0
EATON CORP PLC COMMON STOCK G29183103 22,965 276,189 SH   DFND 1 269,517 0 6,672
EATON CORP PLC COMMON STOCK G29183103 5,215 62,716 SH   OTR 1 58,289 1,465 2,962
EATON CORPORATION PLC COMMON STOCK G29183103 103 1,242 SH   DFND 5 1,242 0 0
ENDO INTERNATIONAL PLC COMMON STOCK G30401106 5 1,521 SH   DFND 1 1,102 0 419
ENDO INTERNATIONAL PLC COMMON STOCK G30401106 5 1,527 SH   OTR 1 0 1,527 0
ENSTAR GROUP LTD COMMON STOCK G3075P101 26 136 SH   DFND 1 19 0 117
ASA GOLD AND PRECIOUS METALS L OTHER G3156P103 0 21 SH   DFND 1 21 0 0
ESSENT GROUP LTD COMMON STOCK G3198U102 149 3,121 SH   DFND 1 3,121 0 0
ESSENT GROUP LTD COMMON STOCK G3198U102 43 892 SH   OTR 1 41 851 0
EVEREST RE GROUP LTD COMMON STOCK G3223R108 2,946 11,074 SH   DFND 1 4,195 0 6,879
EVEREST RE GROUP LTD COMMON STOCK G3223R108 318 1,198 SH   OTR 1 796 5 397
EVEREST RE GROUP LTD COMMON STOCK G3223R108 11 44 SH   DFND 5 44 0 0
FABRINET COMMON STOCK G3323L100 480 9,181 SH   DFND 1 9,166 0 15
FABRINET COMMON STOCK G3323L100 33 634 SH   OTR 1 634 0 0
FERROGLOBE PLC COMMON STOCK G33856108 1 1,128 SH   DFND 1 1,128 0 0
FRESH DEL MONTE PRODUCE INC COMMON STOCK G36738105 442 12,965 SH   DFND 1 12,965 0 0
FRESH DEL MONTE PRODUCE INC COMMON STOCK G36738105 169 4,949 SH   OTR 1 4,949 0 0
EROS INTERNATIONAL PLC COMMON STOCK G3788M114 4 2,183 SH   DFND 1 2,183 0 0
GEOPARK LTD COMMON STOCK G38327105 4,463 242,428 SH   DFND 1 242,428 0 0
GEOPARK LTD COMMON STOCK G38327105 208 11,320 SH   OTR 1 11,320 0 0
GATES INDUSTRIAL CORP PLC COMMON STOCK G39108108 4 380 SH   DFND 1 380 0 0
GENPACT LIMITED COMMON STOCK G3922B107 5,292 136,571 SH   DFND 1 132,275 0 4,296
GENPACT LIMITED COMMON STOCK G3922B107 526 13,574 SH   OTR 1 12,074 1,500 0
GENPACT LTD COMMON STOCK G3922B107 14 379 SH   DFND 5 379 0 0
GLOBAL INDEMNITY LTD COMMON STOCK G3933F105 7 284 SH   DFND 1 284 0 0
GREENLIGHT CAPITAL RE LTD A COMMON STOCK G4095J109 2 234 SH   DFND 1 234 0 0
HELEN OF TROY CORP LTD COMMON STOCK G4388N106 1,301 8,251 SH   DFND 1 8,219 0 32
HELEN OF TROY CORP LTD COMMON STOCK G4388N106 162 1,032 SH   OTR 1 516 516 0
HELEN OF TROY LTD COMMON STOCK G4388N106 1 9 SH   DFND 5 9 0 0
HERBALIFE NUTRITION LTD COMMON STOCK G4412G101 51 1,341 SH   DFND 1 1,150 0 191
HERBALIFE NUTRITION LTD COMMON STOCK G4412G101 11 278 SH   OTR 1 0 0 278
JANUS HENDERSON GROUP PLC COMMON STOCK G4474Y214 78 3,473 SH   DFND 1 1,223 0 2,250
HOLLYSYS AUTOMATION TECHNOLOGI COMMON STOCK G45667105 42 2,791 SH   DFND 1 2,736 0 55
HOLLYSYS AUTOMATION TECHNOLOGI COMMON STOCK G45667105 5 353 SH   OTR 1 353 0 0
HORIZON THERAPEUTICS PLC COMMON STOCK G46188101 996 36,582 SH   DFND 1 35,544 0 1,038
HORIZON THERAPEUTICS PLC COMMON STOCK G46188101 176 6,458 SH   OTR 1 6,118 340 0
HORIZON THERAPEUTICS PLC COMMON STOCK G46188101 1 70 SH   DFND 5 70 0 0
ICON PLC COMMON STOCK G4705A100 703 4,773 SH   DFND 1 4,689 0 84
ICON PLC COMMON STOCK G4705A100 9 64 SH   OTR 1 40 24 0
ICON PLC COMMON STOCK G4705A100 169 1,149 SH   DFND 5 1,149 0 0
ICHOR HOLDINGS INC COMMON STOCK G4740B105 8 328 SH   OTR 1 0 0 328
IHS MARKIT LTD COMMON STOCK G47567105 3,008 44,980 SH   DFND 1 43,389 0 1,591
IHS MARKIT LTD COMMON STOCK G47567105 525 7,851 SH   OTR 1 6,855 757 239
IHS MARKIT LTD COMMON STOCK G47567105 222 3,320 SH   DFND 5 3,320 0 0
INGERSOLL RAND PLC COMMON STOCK G47791101 13,612 110,478 SH   DFND 1 107,668 0 2,810
INGERSOLL RAND PLC COMMON STOCK G47791101 3,556 28,862 SH   OTR 1 24,026 1,322 3,514
INGERSOLL RAND PLC COMMON STOCK G47791101 97 788 SH   DFND 5 788 0 0
INTERNATIONAL GAME TECHNOLOG COMMON STOCK G4863A108 9 629 SH   OTR 1 629 0 0
INVESCO LTD COMMON STOCK G491BT108 280 16,524 SH   DFND 1 11,673 0 4,851
INVESCO LTD COMMON STOCK G491BT108 17 1,011 SH   OTR 1 1,011 0 0
INVESCO LTD COMMON STOCK G491BT108 139 8,259 SH   DFND 5 8,259 0 0
JAMES RIVER GROUP HOLDINGS L COMMON STOCK G5005R107 3 59 SH   DFND 1 59 0 0
JAMES RIVER GROUP HOLDINGS L COMMON STOCK G5005R107 52 1,012 SH   OTR 1 0 1,012 0
JAMES RIVER GROUP HOLDINGS LTD COMMON STOCK G5005R107 50 986 SH   DFND 5 986 0 0
JAZZ PHARMACEUTICALS PLC COMMON STOCK G50871105 1,423 11,106 SH   DFND 1 11,096 0 10
JAZZ PHARMACEUTICALS PLC COMMON STOCK G50871105 728 5,684 SH   OTR 1 3,579 98 2,007
JAZZ PHARMACEUTICALS PLC COMMON STOCK G50871105 71 555 SH   DFND 5 555 0 0
JOHNSON CTLS INTL PLC COMMON STOCK G51502105 1,444 32,886 SH   DFND 1 29,915 0 2,971
JOHNSON CTLS INTL PLC COMMON STOCK G51502105 487 11,085 SH   OTR 1 9,957 418 710
JOHNSON CONTROLS INTERNATIONAL COMMON STOCK G51502105 57 1,305 SH   DFND 5 853 0 452
LAZARD LTD CL A COMMON STOCK G54050102 64 1,818 SH   DFND 1 1,518 0 300
LIBERTY GLOBAL PLC A COMMON STOCK G5480U104 4 177 SH   DFND 1 177 0 0
LIBERTY GLOBAL PLC SERIES C COMMON STOCK G5480U120 636 26,736 SH   DFND 1 26,736 0 0
LIBERTY GLOBAL PLC SERIES C COMMON STOCK G5480U120 87 3,648 SH   OTR 1 3,648 0 0
LIBERTY GLOBAL PLC COMMON STOCK G5480U120 10 445 SH   DFND 5 445 0 0
LINDE PLC COMMON STOCK G5494J103 27,252 140,679 SH   DFND 1 137,409 0 3,270
LINDE PLC COMMON STOCK G5494J103 13,277 68,535 SH   OTR 1 58,905 1,762 7,868
LINDE PLC COMMON STOCK G5494J103 206 1,062 SH   DFND 5 1,062 0 0
LIVANOVA PLC COMMON STOCK G5509L101 41 556 SH   DFND 1 479 0 77
LIVANOVA PLC COMMON STOCK G5509L101 68 918 SH   OTR 1 0 918 0
MALLINCKRODT PLC COMMON STOCK G5785G107 3 1,112 SH   DFND 1 1,112 0 0
MALLINCKRODT PLC COMMON STOCK G5785G107 1 273 SH   OTR 1 242 31 0
MARVELL TECHNOLOGY GROUP LTD COMMON STOCK G5876H105 10,416 417,150 SH   DFND 1 414,909 0 2,241
MARVELL TECHNOLOGY GROUP LTD COMMON STOCK G5876H105 1,201 48,100 SH   OTR 1 47,906 194 0
MARVELL TECHNOLOGY GROUP LTD COMMON STOCK G5876H105 2 89 SH   DFND 5 89 0 0
MEDTRONIC PLC COMMON STOCK G5960L103 65,321 601,369 SH   DFND 1 583,458 0 17,911
MEDTRONIC PLC COMMON STOCK G5960L103 38,671 356,011 SH   OTR 1 312,053 9,593 34,365
MEDTRONIC PLC COMMON STOCK G5960L103 436 4,016 SH   DFND 5 4,016 0 0
APTIV PLC COMMON STOCK G6095L109 19,090 218,376 SH   DFND 1 214,546 0 3,830
APTIV PLC COMMON STOCK G6095L109 2,684 30,704 SH   OTR 1 29,482 407 815
APTIV PLC COMMON STOCK G6095L109 10 116 SH   DFND 5 116 0 0
NABORS INDUSTRIES LTD COMMON STOCK G6359F103 6 3,160 SH   DFND 1 3,160 0 0
NABORS INDUSTRIES LTD COMMON STOCK G6359F103 3 1,552 SH   OTR 1 0 1,552 0
NABRIVA THERAPEUTICS PLC COMMON STOCK G63637105 1 500 SH   DFND 1 500 0 0
NAM TAI PROPERTY INC COMMON STOCK G63907102 1 165 SH   OTR 1 165 0 0
NIELSEN HOLDINGS PLC COMMON STOCK G6518L108 69 3,236 SH   DFND 1 2,706 0 530
NOBLE CORP PLC COMMON STOCK G65431101 1 931 SH   DFND 1 931 0 0
NOMAD FOODS LTD COMMON STOCK G6564A105 2,365 115,419 SH   DFND 1 115,302 0 117
NOMAD FOODS LTD COMMON STOCK G6564A105 403 19,659 SH   OTR 1 19,659 0 0
NORWEGIAN CRUISE LINE HOLDINGS COMMON STOCK G66721104 643 12,416 SH   DFND 1 12,059 0 357
NORWEGIAN CRUISE LINE HOLDINGS COMMON STOCK G66721104 100 1,927 SH   OTR 1 1,162 765 0
NOVOCURE LTD COMMON STOCK G6674U108 793 10,600 SH   DFND 1 10,579 0 21
NOVOCURE LTD COMMON STOCK G6674U108 23 311 SH   OTR 1 311 0 0
NVENT ELECTRIC PLC COMMON STOCK G6700G107 6,694 303,722 SH   DFND 1 302,206 0 1,516
NVENT ELECTRIC PLC COMMON STOCK G6700G107 13,275 602,343 SH   OTR 1 588,113 104 14,126
PAGSEGURO DIGITAL LTD CL A COMMON STOCK G68707101 9 200 SH   DFND 1 200 0 0
PAGSEGURO DIGITAL LTD CL A COMMON STOCK G68707101 23 500 SH   OTR 1 500 0 0
PROTHENA CORP PLC COMMON STOCK G72800108 9 1,160 SH   DFND 1 1,160 0 0
RENAISSANCE RE HOLDINGS LTD COMMON STOCK G7496G103 1,330 6,874 SH   DFND 1 6,763 0 111
RENAISSANCE RE HOLDINGS LTD COMMON STOCK G7496G103 241 1,246 SH   OTR 1 1,139 0 107
RENAISSANCERE HOLDINGS LTD COMMON STOCK G7496G103 24 128 SH   DFND 5 128 0 0
SEAGATE TECHNOLOGY COMMON STOCK G7945M107 389 7,228 SH   DFND 1 5,963 0 1,265
SEAGATE TECHNOLOGY COMMON STOCK G7945M107 107 1,991 SH   OTR 1 1,541 0 450
SEADRILL LTD COMMON STOCK G7998G106 0 3 SH   DFND 1 3 0 0
PENTAIR PLC COMMON STOCK G7S00T104 10,550 279,091 SH   DFND 1 277,789 0 1,302
PENTAIR PLC COMMON STOCK G7S00T104 21,496 568,670 SH   OTR 1 552,793 2,178 13,699
SENSATA TECHNOLOGIES HOLDING COMMON STOCK G8060N102 1,643 32,808 SH   DFND 1 32,247 0 561
SENSATA TECHNOLOGIES HOLDING COMMON STOCK G8060N102 288 5,754 SH   OTR 1 5,754 0 0
SENSATA TECHNOLOGIES HLDG PLC COMMON STOCK G8060N102 29 589 SH   DFND 5 589 0 0
SHIP FINANCE INTERNATIONAL LTD COMMON STOCK G81075106 53 3,740 SH   DFND 1 3,740 0 0
SIGNET JEWELERS LTD COMMON STOCK G81276100 532 31,714 SH   DFND 1 205 0 31,509
SIGNET JEWELERS LTD COMMON STOCK G81276100 4 243 SH   OTR 1 0 243 0
SINA CORP COMMON STOCK G81477104 139 3,556 SH   DFND 1 2,600 0 956
SINA CORP COMMON STOCK G81477104 7 194 SH   OTR 1 165 0 29
SOCIAL CAPITAL HEDOSOPHIA HO COMMON STOCK G8250R103 5 500 SH   DFND 1 500 0 0
STERIS PLC COMMON STOCK G8473T100 5,489 37,986 SH   DFND 1 35,225 0 2,761
STERIS PLC COMMON STOCK G8473T100 897 6,213 SH   OTR 1 5,046 0 1,167
STERIS PLC COMMON STOCK G8473T100 547 3,787 SH   DFND 5 3,787 0 0
STONECO LTD A COMMON STOCK G85158106 122 3,503 SH   DFND 1 3,503 0 0
STONECO LTD A COMMON STOCK G85158106 104 3,000 SH   OTR 1 3,000 0 0
TECHNIPFMC PLC COMMON STOCK G87110105 303 12,555 SH   DFND 1 9,106 0 3,449
TECHNIPFMC PLC COMMON STOCK G87110105 15 610 SH   OTR 1 481 0 129
TECHNIPFMC PLC COMMON STOCK G87110105 34 1,422 SH   DFND 5 1,422 0 0
TEXTAINER GROUP HOLDINGS LTD COMMON STOCK G8766E109 738 74,458 SH   DFND 1 0 0 74,458
THERAVANCE BIOPHARMA INC COMMON STOCK G8807B106 1 75 SH   DFND 1 0 0 75
LIBERTY LATIN AMERICA LTD COMMON STOCK G9001E102 17 1,000 SH   DFND 1 1,000 0 0
LIBERTY LATIN AMERICA LTD COMMON STOCK G9001E128 3 174 SH   DFND 1 174 0 0
LIBERTY LATIN AMERICA LTD COMMON STOCK G9001E128 45 2,649 SH   OTR 1 0 2,649 0
TRITON INTERNATIONAL LTD BER COMMON STOCK G9078F107 5 141 SH   DFND 1 141 0 0
TRITON INTERNATIONAL LTD BER COMMON STOCK G9078F107 2 68 SH   OTR 1 0 68 0
TRONOX HOLDINGS PLC CLASS A COMMON STOCK G9087Q102 3 375 SH   DFND 1 258 0 117
TSAKOS ENERGY NAVIGATION LTD COMMON STOCK G9108L108 177 60,000 SH   DFND 1 60,000 0 0
VALARIS PLC COMMON STOCK G9402V109 32 6,674 SH   DFND 1 5,882 0 792
VALARIS PLC COMMON STOCK G9402V109 1 258 SH   OTR 1 0 0 258
GOLAR LNG LTD COMMON STOCK G9456A100 5 380 SH   DFND 1 380 0 0
WHITE MTNS INS GROUP LTD COMMON STOCK G9618E107 85 79 SH   DFND 1 51 0 28
WHITE MTNS INS GROUP LTD COMMON STOCK G9618E107 205 190 SH   OTR 1 114 72 4
WHITE MOUNTAINS INSURANCE GP COMMON STOCK G9618E107 63 58 SH   DFND 5 58 0 0
WILLIS TOWERS WATSON PLC COMMON STOCK G96629103 6,439 33,370 SH   DFND 1 32,888 0 482
WILLIS TOWERS WATSON PLC COMMON STOCK G96629103 794 4,114 SH   OTR 1 3,862 233 19
PERRIGO CO PLC COMMON STOCK G97822103 826 14,780 SH   DFND 1 14,479 0 301
PERRIGO CO PLC COMMON STOCK G97822103 12 219 SH   OTR 1 105 86 28
ALCON INC COMMON STOCK H01301128 4,357 74,746 SH   DFND 1 71,092 0 3,654
ALCON INC COMMON STOCK H01301128 845 14,494 SH   OTR 1 11,171 2,348 975
ALCON INC COMMON STOCK H01301128 385 6,616 SH   DFND 5 6,616 0 0
CHUBB LTD COMMON STOCK H1467J104 127,957 792,595 SH   DFND 1 784,467 0 8,128
CHUBB LTD COMMON STOCK H1467J104 31,009 192,081 SH   OTR 1 179,195 6,052 6,834
CHUBB LTD COMMON STOCK H1467J104 715 4,431 SH   DFND 5 4,431 0 0
CRISPR THERAPEUTICS AG COMMON STOCK H17182108 3 75 SH   DFND 1 75 0 0
CRISPR THERAPEUTICS AG COMMON STOCK H17182108 4 95 SH   OTR 1 95 0 0
GARMIN LTD COMMON STOCK H2906T109 312 3,679 SH   DFND 1 3,439 0 240
GARMIN LTD COMMON STOCK H2906T109 48 557 SH   OTR 1 443 44 70
GARMIN LTD COMMON STOCK H2906T109 8 104 SH   DFND 5 104 0 0
UBS GROUP AG COMMON STOCK H42097107 334 29,517 SH   DFND 1 21,717 0 7,800
UBS GROUP AG COMMON STOCK H42097107 85 7,484 SH   OTR 1 3,755 955 2,774
UBS GROUP AG COMMON STOCK H42097107 4 425 SH   DFND 5 425 0 0
LOGITECH INTL SA COMMON STOCK H50430232 484 11,892 SH   DFND 1 11,868 0 24
LOGITECH INTL SA COMMON STOCK H50430232 46 1,106 SH   OTR 1 1,106 0 0
OBSEVA SA COMMON STOCK H5861P103 108 13,000 SH   OTR 1 13,000 0 0
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GLOBANT SA COMMON STOCK L44385109 120 1,311 SH   OTR 1 1,311 0 0
GLOBANT S A COMMON STOCK L44385109 22 247 SH   DFND 5 247 0 0
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MILLICOM INTL CELLULAR SA COMMON STOCK L6388F110 1 12 SH   DFND 1 12 0 0
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ORION ENGINEERED CARBONS SA COMMON STOCK L72967109 3 183 SH   OTR 1 0 183 0
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TRINSEO SA COMMON STOCK L9340P101 3 60 SH   OTR 1 0 0 60
AUDIOCODES LTD COMMON STOCK M15342104 2 149 SH   DFND 5 149 0 0
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CHECK POINT SOFTWARE TECH LTD COMMON STOCK M22465104 2,942 26,873 SH   OTR 1 22,185 1,279 3,409
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CYBER ARK SOFTWARE LTD COMMON STOCK M2682V108 187 1,878 SH   OTR 1 1,801 77 0
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GALMED PHARMACEUTICALS LTD COMMON STOCK M47238106 9 1,880 SH   OTR 1 0 0 1,880
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MELLANOX TECHNOLOGIES LTD COMMON STOCK M51363113 42 385 SH   OTR 1 265 120 0
MELLANOX TECHNOLOGIES INC COMMON STOCK M51363113 1 18 SH   DFND 5 18 0 0
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WIX COM LTD COMMON STOCK M98068105 13 111 SH   DFND 1 111 0 0
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FERRARI NV COMMON STOCK N3167Y103 6 41 SH   DFND 5 41 0 0
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AKEBIA THERAPEUTICS INC COMMON STOCK 00972D105 8 2,000 SH   OTR 1 2,000 0 0
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ALARM COM HOLDINGS INC COMMON STOCK 011642105 117 2,505 SH   OTR 1 2,505 0 0
ALARM.COM INC COMMON STOCK 011642105 18 404 SH   DFND 5 404 0 0
ALASKA AIR GROUP INC COMMON STOCK 011659109 482 7,430 SH   DFND 1 7,253 0 177
ALASKA AIR GROUP INC COMMON STOCK 011659109 107 1,646 SH   OTR 1 846 800 0
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ALBANY INTL CORP CL A COMMON STOCK 012348108 206 2,290 SH   OTR 1 31 909 1,350
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ALBEMARLE CORP COMMON STOCK 012653101 4,897 70,440 SH   OTR 1 69,086 1,325 29
ALBEMARLE CORP COMMON STOCK 012653101 17 256 SH   DFND 5 256 0 0
ALCOA CORPORATION COMMON STOCK 013872106 126 6,279 SH   DFND 1 6,114 0 165
ALCOA CORPORATION COMMON STOCK 013872106 32 1,628 SH   OTR 1 1,612 0 16
ALDER BIOPHARMACEUTICALS INC COMMON STOCK 014339105 19 1,000 SH   DFND 1 1,000 0 0
ALECTOR INC COMMON STOCK 014442107 4 250 SH   DFND 1 250 0 0
ALERUS FINANCIAL CORP COMMON STOCK 01446U103 370 16,955 SH   DFND 1 16,955 0 0
ALEXANDER & BALDWIN INC COMMON STOCK 014491104 14 566 SH   DFND 1 414 0 152
ALEXANDER & BALDWIN INC COMMON STOCK 014491104 3 116 SH   OTR 1 0 0 116
ALEXANDERS INC COMMON STOCK 014752109 1 2 SH   DFND 1 2 0 0
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ALEXANDRIA REAL ESTATE EQUITIE COMMON STOCK 015271109 495 3,213 SH   OTR 1 2,813 382 18
ALEXANDRIA REAL ESTATE EQUITY COMMON STOCK 015271109 31 202 SH   DFND 5 202 0 0
ALEXION PHARMACEUTICALS INC COMMON STOCK 015351109 2,545 25,983 SH   DFND 1 25,412 0 571
ALEXION PHARMACEUTICALS INC COMMON STOCK 015351109 754 7,696 SH   OTR 1 3,078 4,287 331
ALEXION PHARMACEUTICALS INC COMMON STOCK 015351109 185 1,898 SH   DFND 5 1,898 0 0
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ALIBABA GROUP HOLDING LTD A D COMMON STOCK 01609W102 2,929 17,514 SH   OTR 1 8,807 3,235 5,472
ALIBABA GROUP HOLDING LTD COMMON STOCK 01609W102 198 1,187 SH   DFND 5 1,084 0 103
ALICO INC COMMON STOCK 016230104 32 933 SH   DFND 1 933 0 0
ALIGN TECHNOLOGY INC COMMON STOCK 016255101 19,478 107,664 SH   DFND 1 107,537 0 127
ALIGN TECHNOLOGY INC COMMON STOCK 016255101 6,587 36,408 SH   OTR 1 34,138 1,470 800
ALIGN TECHNOLOGY INC COMMON STOCK 016255101 14 79 SH   DFND 5 79 0 0
ALLEGHANY CORP COMMON STOCK 017175100 51 64 SH   DFND 1 64 0 0
ALLEGHENY TECHNOLOGIES INC COMMON STOCK 01741R102 784 38,708 SH   DFND 1 33,957 0 4,751
ALLEGHENY TECHNOLOGIES INC COMMON STOCK 01741R102 152 7,522 SH   OTR 1 7,376 0 146
ALLEGHENY TECH INC COMMON STOCK 01741R102 15 785 SH   DFND 5 785 0 0
ALLEGIANT TRAVEL CO COMMON STOCK 01748X102 132 884 SH   DFND 1 21 0 863
ALLETE INC COMMON STOCK 018522300 1,142 13,067 SH   DFND 1 11,857 0 1,210
ALLETE INC COMMON STOCK 018522300 920 10,531 SH   OTR 1 8,831 400 1,300
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ALLIANCE DATA SYSTEMS CORP COMMON STOCK 018581108 16 126 SH   OTR 1 100 0 26
ALLIANCEBERNSTEIN NATIONAL MUN OTHER 01864U106 176 12,787 SH   OTR 1 12,787 0 0
ALLIANCE RESOURCE PARTNERS L P COMMON STOCK 01877R108 53 3,300 SH   DFND 1 3,300 0 0
ALLIANCEBERNSTEIN GL HIGH INCO OTHER 01879R106 78 6,530 SH   DFND 1 6,530 0 0
ALLIANCEBERNSTEIN GL HIGH INCO OTHER 01879R106 107 8,933 SH   OTR 1 5,921 0 3,012
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ALLIANT ENERGY CORP COMMON STOCK 018802108 6,072 112,578 SH   DFND 1 106,061 0 6,517
ALLIANT ENERGY CORP COMMON STOCK 018802108 3,342 61,975 SH   OTR 1 59,333 2,642 0
ALLIANT ENERGY CORP COMMON STOCK 018802108 7 144 SH   DFND 5 0 0 144
ALLIANCEBERNSTEIN HLDG LP COMMON STOCK 01881G106 16 550 SH   DFND 1 550 0 0
ALLIANCEBERNSTEIN HLDG LP COMMON STOCK 01881G106 183 6,250 SH   OTR 1 4,000 0 2,250
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ALLIANZGI NFJ DIVIDEND INTER OTHER 01883A107 118 9,629 SH   DFND 1 9,629 0 0
ALLIANZGI NFJ DIVIDEND INTER OTHER 01883A107 34 2,775 SH   OTR 1 2,775 0 0
ALLIED MOTION TECHNOLOGIES INC COMMON STOCK 019330109 5 135 SH   OTR 1 135 0 0
ALLISON TRANSMISSION HOLDINGS COMMON STOCK 01973R101 411 8,733 SH   DFND 1 8,624 0 109
ALLISON TRANSMISSION HOLDINGS COMMON STOCK 01973R101 159 3,377 SH   OTR 1 1,657 1,720 0
ALLISON TRANSMISSION HLGDS INC COMMON STOCK 01973R101 3 68 SH   DFND 5 68 0 0
ALLOGENE THERAPEUTICS INC COMMON STOCK 019770106 1 50 SH   OTR 1 50 0 0
ALLSTATE CORP COMMON STOCK 020002101 13,166 121,147 SH   DFND 1 118,049 0 3,098
ALLSTATE CORP COMMON STOCK 020002101 7,875 72,460 SH   OTR 1 70,619 402 1,439
ALLSTATE CORP COMMON STOCK 020002101 128 1,185 SH   DFND 5 1,185 0 0
ALLY FINANCIAL INC COMMON STOCK 02005N100 362 10,905 SH   DFND 1 7,539 0 3,366
ALLY FINANCIAL INC COMMON STOCK 02005N100 27 821 SH   OTR 1 0 0 821
ALLY FINANCIAL INC COMMON STOCK 02005N100 39 1,187 SH   DFND 5 1,187 0 0
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ALNYLAM PHARMACEUTICALS INC COMMON STOCK 02043Q107 17 216 SH   OTR 1 0 112 104
ALPHABET INC CL C COMMON STOCK 02079K107 116,908 95,905 SH   DFND 1 90,132 0 5,773
ALPHABET INC CL C COMMON STOCK 02079K107 36,315 29,791 SH   OTR 1 24,737 1,742 3,312
ALPHABET INC COMMON STOCK 02079K107 804 660 SH   DFND 5 660 0 0
ALPHABET INC CL A COMMON STOCK 02079K305 250,083 204,795 SH   DFND 1 200,326 0 4,469
ALPHABET INC CL A COMMON STOCK 02079K305 60,294 49,375 SH   OTR 1 42,450 2,719 4,206
ALPHABET INC COMMON STOCK 02079K305 1,033 846 SH   DFND 5 846 0 0
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ALTABA INC OTHER 021346101 185 9,482 SH   OTR 1 9,482 0 0
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ALTAIR ENGINEERING INC A C COMMON STOCK 021369103 140 4,045 SH   OTR 1 0 1,495 2,550
ALTERYX INC CLASS A COMMON STOCK 02156B103 68 636 SH   OTR 1 200 377 59
ALTICE USA INC A COMMON STOCK 02156K103 388 13,545 SH   DFND 1 13,545 0 0
ALTICE USA INC A COMMON STOCK 02156K103 69 2,385 SH   OTR 1 2,385 0 0
ALTICE USA INC COMMON STOCK 02156K103 8 295 SH   DFND 5 295 0 0
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ALTRA INDUSTRIAL MOTION CORP COMMON STOCK 02208R106 8 278 SH   OTR 1 278 0 0
ALTRIA GROUP INC COMMON STOCK 02209S103 22,691 554,788 SH   DFND 1 537,508 0 17,280
ALTRIA GROUP INC COMMON STOCK 02209S103 18,138 443,476 SH   OTR 1 430,585 5,270 7,621
ALTRIA GROUP INC COMMON STOCK 02209S103 268 6,560 SH   DFND 5 6,428 0 132
ALTUS MIDSTREAM CO COMMON STOCK 02215L100 1 193 SH   DFND 5 193 0 0
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ALUMINUM CORP CHINA A D R COMMON STOCK 022276109 1 110 SH   OTR 1 110 0 0
AMALGAMATED BK OF NEW YORK COMMON STOCK 022663108 50 3,140 SH   DFND 1 3,140 0 0
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AMAZON COM INC COMMON STOCK 023135106 92,037 53,019 SH   OTR 1 44,091 2,395 6,533
AMAZON.COM INC COMMON STOCK 023135106 1,197 690 SH   DFND 5 678 0 12
AMBAC FINANCIAL GROUP INC COMMON STOCK 023139884 9 461 SH   DFND 1 461 0 0
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AMBEV SA COMMON STOCK 02319V103 15 3,395 SH   DFND 5 3,395 0 0
AMEDISYS INC COMMON STOCK 023436108 1,293 9,865 SH   DFND 1 9,795 0 70
AMEDISYS INC COMMON STOCK 023436108 388 2,961 SH   OTR 1 2,691 270 0
AMEDISYS INC COMMON STOCK 023436108 17 132 SH   DFND 5 132 0 0
AMERCO COMMON STOCK 023586100 14 37 SH   DFND 1 37 0 0
AMERCO COMMON STOCK 023586100 1 3 SH   OTR 1 0 0 3
AMERCO COMMON STOCK 023586100 49 127 SH   DFND 5 127 0 0
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AMEREN CORP COMMON STOCK 023608102 2,291 28,614 SH   OTR 1 21,867 3,047 3,700
AMEREN CORP COMMON STOCK 023608102 36 456 SH   DFND 5 456 0 0
AMERICA FIRST MULTIFAMILY IN L COMMON STOCK 02364V107 2,435 297,703 SH   DFND 1 297,703 0 0
AMERICA MOVIL A D R COMMON STOCK 02364W105 297 19,994 SH   DFND 1 19,455 0 539
AMERICA MOVIL A D R COMMON STOCK 02364W105 1,325 89,195 SH   OTR 1 85,475 2,997 723
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AMERICAN AIRLINES GROUP INC COMMON STOCK 02376R102 1 22 SH   OTR 1 22 0 0
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AMERICAN AXLE & MFG HLDGS INC COMMON STOCK 024061103 68 8,261 SH   DFND 1 8,261 0 0
AMERICAN AXLE & MFG HLDGS INC COMMON STOCK 024061103 5 565 SH   OTR 1 565 0 0
AMERICAN CAMPUS COMMUNITIES COMMON STOCK 024835100 44,190 919,088 SH   DFND 1 914,341 0 4,747
AMERICAN CAMPUS COMMUNITIES COMMON STOCK 024835100 6,850 142,449 SH   OTR 1 141,582 98 769
AMERICAN CAMPUS COMMUNITIES COMMON STOCK 024835100 72 1,518 SH   DFND 5 1,518 0 0
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AMERICAN ELECTRIC POWER COMMON STOCK 025537101 9,288 99,128 SH   OTR 1 95,316 625 3,187
AMERICAN ELECTRIC POWER CO INC COMMON STOCK 025537101 53 573 SH   DFND 5 573 0 0
AMERICAN EAGLE OUTFITTERS COMMON STOCK 02553E106 41 2,503 SH   DFND 1 2,503 0 0
AMERICAN EQUITY INVT LIFE HL COMMON STOCK 025676206 655 27,074 SH   DFND 1 25,005 0 2,069
AMERICAN EQUITY INVT LIFE HL COMMON STOCK 025676206 101 4,173 SH   OTR 1 4,173 0 0
AMERICAN EXPRESS CO COMMON STOCK 025816109 33,315 281,663 SH   DFND 1 273,519 0 8,144
AMERICAN EXPRESS CO COMMON STOCK 025816109 20,574 173,953 SH   OTR 1 164,629 1,301 8,023
AMERICAN EXPRESS COMPANY COMMON STOCK 025816109 281 2,383 SH   DFND 5 2,383 0 0
AMERICAN FINL GROUP INC OHIO COMMON STOCK 025932104 382 3,543 SH   DFND 1 2,867 0 676
AMERICAN FINL GROUP INC OHIO COMMON STOCK 025932104 78 726 SH   OTR 1 533 0 193
AMERICAN HOMES 4 RENT A COMMON STOCK 02665T306 753 29,088 SH   DFND 1 28,935 0 153
AMERICAN HOMES 4 RENT A COMMON STOCK 02665T306 25 980 SH   OTR 1 907 0 73
AMERICAN INTL GROUP INC WT 1 OTHER 026874156 8 539 SH   DFND 1 539 0 0
AMERICAN INTL GROUP INC WT 1 OTHER 026874156 0 18 SH   OTR 1 18 0 0
AMERICAN INTERNATIONAL GROUP COMMON STOCK 026874784 2,960 53,137 SH   DFND 1 47,921 0 5,216
AMERICAN INTERNATIONAL GROUP COMMON STOCK 026874784 430 7,724 SH   OTR 1 7,045 161 518
AMERICAN INTERNATIONAL GROUP COMMON STOCK 026874784 77 1,384 SH   DFND 5 1,384 0 0
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AMERICAN NATL INS CO COMMON STOCK 028591105 3 22 SH   OTR 1 0 22 0
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AMERICAN STATES WATER CO COMMON STOCK 029899101 289 3,218 SH   DFND 1 2,951 0 267
AMERICAN STATES WATER CO COMMON STOCK 029899101 180 2,000 SH   OTR 1 1,000 0 1,000
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AMERICAN VANGUARD CORP COMMON STOCK 030371108 12 742 SH   DFND 1 742 0 0
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AMERICAN WOODMARK CORP COMMON STOCK 030506109 29 328 SH   OTR 1 328 0 0
AMERICAN WOODMARK CORP COMMON STOCK 030506109 13 153 SH   DFND 5 153 0 0
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AMERISAFE INC COMMON STOCK 03071H100 23 352 SH   OTR 1 0 352 0
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AMERISOURCEBERGEN CORP COMMON STOCK 03073E105 435 5,286 SH   DFND 5 5,286 0 0
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AMERIPRISE FINL INC COMMON STOCK 03076C106 4,122 28,022 SH   OTR 1 27,377 179 466
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AMERIS BANCORP COMMON STOCK 03076K108 56 1,369 SH   OTR 1 0 109 1,260
AMES NATL CORP COMMON STOCK 031001100 305 10,651 SH   DFND 1 10,651 0 0
AMES NATL CORP COMMON STOCK 031001100 63 2,204 SH   OTR 1 2,204 0 0
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AMETEK INC COMMON STOCK 031100100 1,622 17,656 SH   OTR 1 16,717 759 180
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AMGEN INC COMMON STOCK 031162100 102,818 531,330 SH   DFND 1 507,731 0 23,599
AMGEN INC COMMON STOCK 031162100 32,942 170,237 SH   OTR 1 151,206 2,279 16,752
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AMKOR TECHNOLOGY INC COMMON STOCK 031652100 14 1,500 SH   DFND 1 1,500 0 0
AMNEAL PHARMACEUTICALS INC COMMON STOCK 03168L105 2 634 SH   DFND 1 634 0 0
AMPCO-PITTSBURG CORP COMMON STOCK 032037103 3 843 SH   DFND 1 843 0 0
AMPHENOL CORP CL A COMMON STOCK 032095101 18,244 189,065 SH   DFND 1 187,624 0 1,441
AMPHENOL CORP CL A COMMON STOCK 032095101 5,275 54,668 SH   OTR 1 46,382 1,439 6,847
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ANALOG DEVICES INC COMMON STOCK 032654105 3,742 33,486 SH   OTR 1 21,670 2,403 9,413
ANALOG DEVICES INC COMMON STOCK 032654105 45 411 SH   DFND 5 411 0 0
ANAPTYSBIO INC COMMON STOCK 032724106 14 400 SH   OTR 1 400 0 0
ANAPLAN INC COMMON STOCK 03272L108 21 455 SH   DFND 1 455 0 0
ANGIODYNAMICS INC COMMON STOCK 03475V101 87 4,746 SH   DFND 1 4,746 0 0
ANGIODYNAMICS INC COMMON STOCK 03475V101 15 813 SH   OTR 1 813 0 0
ANGIODYNAMICS INC COMMON STOCK 03475V101 6 367 SH   DFND 5 367 0 0
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ANGLOGOLD ASHANTI LTD SPON A D COMMON STOCK 035128206 5 259 SH   OTR 1 259 0 0
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ANHEUSER BUSCH INBEV NV A D R COMMON STOCK 03524A108 17,418 183,060 SH   OTR 1 182,575 342 143
ANHEUSER BUSCH INBEV COMMON STOCK 03524A108 246 2,590 SH   DFND 5 2,590 0 0
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ANNALY CAPITAL MANAGEMENT INC COMMON STOCK 035710409 437 49,604 SH   OTR 1 49,604 0 0
ANNALY CAPITAL MANAGEMENT COMMON STOCK 035710409 3 376 SH   DFND 5 376 0 0
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ANTHEM INC COMMON STOCK 036752103 3,811 15,870 SH   OTR 1 8,626 668 6,576
ANTHEM INC COMMON STOCK 036752103 279 1,163 SH   DFND 5 1,163 0 0
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ANTERO MIDSTREAM CORP COMMON STOCK 03676B102 1 168 SH   DFND 5 168 0 0
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ANWORTH MORTGAGE ASSET CORP COMMON STOCK 037347101 13 3,800 SH   OTR 1 0 0 3,800
APACHE CORP COMMON STOCK 037411105 1,485 58,020 SH   DFND 1 54,726 0 3,294
APACHE CORP COMMON STOCK 037411105 673 26,305 SH   OTR 1 26,145 0 160
APACHE CORPORATION COMMON STOCK 037411105 1 45 SH   DFND 5 45 0 0
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APARTMENT INVT MGMT CO A COMMON STOCK 03748R754 295 5,646 SH   OTR 1 5,612 0 34
APARTMENT INVESTMENT & MGMT CO COMMON STOCK 03748R754 13 263 SH   DFND 5 263 0 0
APERGY CORP COMMON STOCK 03755L104 1,536 56,794 SH   DFND 1 54,964 0 1,830
APERGY CORP COMMON STOCK 03755L104 712 26,338 SH   OTR 1 24,422 0 1,916
APERGY CORP COMMON STOCK 03755L104 7 264 SH   DFND 5 264 0 0
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APOGEE ENTERPRISES INC COMMON STOCK 037598109 149 3,828 SH   OTR 1 3,828 0 0
APOLLO INVESTMENT CORP COMMON STOCK 03761U502 19 1,174 SH   DFND 1 0 0 1,174
APOLLO INVESTMENT CORP COMMON STOCK 03761U502 27 1,666 SH   OTR 1 1,666 0 0
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APHRIA INC COMMON STOCK 03765K104 1 100 SH   DFND 1 100 0 0
APHRIA INC COMMON STOCK 03765K104 5 900 SH   OTR 1 900 0 0
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APPLE INC COMMON STOCK 037833100 549,524 2,453,556 SH   DFND 1 2,388,835 0 64,721
APPLE INC COMMON STOCK 037833100 208,685 931,752 SH   OTR 1 849,564 29,806 52,382
APPLE INC COMMON STOCK 037833100 670 2,993 SH   DFND 5 2,993 0 0
APPLE HOSPITALITY REIT INC COMMON STOCK 03784Y200 243 14,683 SH   DFND 1 14,253 0 430
APPLE HOSPITALITY REIT INC COMMON STOCK 03784Y200 12 707 SH   OTR 1 0 333 374
APPLIED INDL TECHNOLOGIES INC COMMON STOCK 03820C105 3,305 58,195 SH   DFND 1 5,684 0 52,511
APPLIED INDL TECHNOLOGIES INC COMMON STOCK 03820C105 19 336 SH   OTR 1 276 0 60
APPLIED MATERIALS INC COMMON STOCK 038222105 13,732 275,199 SH   DFND 1 265,664 0 9,535
APPLIED MATERIALS INC COMMON STOCK 038222105 4,117 82,490 SH   OTR 1 73,641 2,990 5,859
APPLIED MATERIALS INC COMMON STOCK 038222105 37 756 SH   DFND 5 756 0 0
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CME GROUP INC COMMON STOCK 12572Q105 165 785 SH   DFND 5 280 0 505
C M S ENERGY CORP COMMON STOCK 125896100 9,497 148,517 SH   DFND 1 147,483 0 1,034
C M S ENERGY CORP COMMON STOCK 125896100 5,206 81,402 SH   OTR 1 78,828 133 2,441
CMS ENERGY CORPORATION COMMON STOCK 125896100 34 538 SH   DFND 5 538 0 0
CPI AEROSTRUCTURES INC COMMON STOCK 125919308 6 767 SH   DFND 1 767 0 0
CNA FINANCIAL CORP COMMON STOCK 126117100 49 989 SH   DFND 1 961 0 28
CNA FINANCIAL CORP COMMON STOCK 126117100 38 755 SH   OTR 1 0 49 706
CNOOC LTD A D R COMMON STOCK 126132109 1,065 6,996 SH   DFND 1 6,616 0 380
CNOOC LTD A D R COMMON STOCK 126132109 144 943 SH   OTR 1 930 0 13
CPFL ENERGIA S A A D R COMMON STOCK 126153105 19 1,219 SH   DFND 1 1,028 0 191
CNO FINANCIAL GROUP INC COMMON STOCK 12621E103 140 8,866 SH   DFND 1 7,077 0 1,789
CNO FINANCIAL GROUP INC COMMON STOCK 12621E103 60 3,773 SH   OTR 1 0 3,773 0
CRH SPON A D R COMMON STOCK 12626K203 1,716 49,947 SH   DFND 1 47,639 0 2,308
CRH SPON A D R COMMON STOCK 12626K203 453 13,182 SH   OTR 1 12,872 113 197
CRH PLC COMMON STOCK 12626K203 21 628 SH   DFND 5 628 0 0
CSG SYS INTL INC COMMON STOCK 126349109 38 729 SH   DFND 1 729 0 0
CSG SYS INTL INC COMMON STOCK 126349109 66 1,285 SH   OTR 1 1,285 0 0
CSW INDUSTRIALS INC COMMON STOCK 126402106 296 4,286 SH   DFND 1 4,286 0 0
CSW INDUSTRIALS INC COMMON STOCK 126402106 53 777 SH   OTR 1 777 0 0
CSX CORP COMMON STOCK 126408103 18,463 266,540 SH   DFND 1 260,788 0 5,752
CSX CORP COMMON STOCK 126408103 13,535 195,389 SH   OTR 1 191,392 1,242 2,755
CSX CORP COMMON STOCK 126408103 32 472 SH   DFND 5 472 0 0
CTS CORP COMMON STOCK 126501105 187 5,755 SH   DFND 1 5,755 0 0
CTS CORP COMMON STOCK 126501105 20 610 SH   OTR 1 610 0 0
CTS CORP COMMON STOCK 126501105 16 502 SH   DFND 5 502 0 0
CNX RESOURCES CORP COMMON STOCK 12653C108 35 4,856 SH   DFND 1 3,168 0 1,688
CNX RESOURCES CORP COMMON STOCK 12653C108 1 195 SH   OTR 1 0 0 195
CUI GLOBAL INC COMMON STOCK 126576206 78 91,890 SH   DFND 1 0 0 91,890
CVB FINL CORP COMMON STOCK 126600105 472 22,609 SH   DFND 1 22,609 0 0
CVB FINL CORP COMMON STOCK 126600105 107 5,146 SH   OTR 1 5,146 0 0
CVR ENERGY INC COMMON STOCK 12662P108 431 9,794 SH   DFND 1 9,794 0 0
CVR ENERGY INC COMMON STOCK 12662P108 45 1,018 SH   OTR 1 897 121 0
CVS HEALTH CORP COMMON STOCK 126650100 34,386 545,195 SH   DFND 1 529,590 0 15,605
CVS HEALTH CORP COMMON STOCK 126650100 10,570 167,599 SH   OTR 1 151,631 5,121 10,847
CVS HEALTH CORP COMMON STOCK 126650100 153 2,430 SH   DFND 5 2,430 0 0
C J ENERGY SVCS INC COMMON STOCK 12674R100 7 690 SH   DFND 1 495 0 195
C J ENERGY SVCS INC COMMON STOCK 12674R100 5 453 SH   OTR 1 0 453 0
CABLE ONE INC COMMON STOCK 12685J105 310 247 SH   DFND 1 219 0 28
CABLE ONE INC COMMON STOCK 12685J105 113 90 SH   OTR 1 0 0 90
CABLE ONE INC COMMON STOCK 12685J105 30 24 SH   DFND 5 24 0 0
CABOT CORP COMMON STOCK 127055101 1,092 24,094 SH   DFND 1 22,479 0 1,615
CABOT CORP COMMON STOCK 127055101 74 1,625 SH   OTR 1 1,625 0 0
CABOT CORPORATION COMMON STOCK 127055101 7 155 SH   DFND 5 155 0 0
CABOT OIL GAS CORP CL A COMMON STOCK 127097103 11,905 677,533 SH   DFND 1 670,705 0 6,828
CABOT OIL GAS CORP CL A COMMON STOCK 127097103 1,501 85,437 SH   OTR 1 82,952 660 1,825
CABOT MICROELECTRONICS CORPORA COMMON STOCK 12709P103 1,477 10,461 SH   DFND 1 10,461 0 0
CABOT MICROELECTRONICS CORPORA COMMON STOCK 12709P103 196 1,387 SH   OTR 1 1,387 0 0
CACI INTERNATIONAL INC CL A COMMON STOCK 127190304 2,670 11,547 SH   DFND 1 10,628 0 919
CACI INTERNATIONAL INC CL A COMMON STOCK 127190304 664 2,873 SH   OTR 1 2,155 679 39
CACI INTERNATIONAL INC COMMON STOCK 127190304 25 110 SH   DFND 5 110 0 0
CACTUS INC A COMMON STOCK 127203107 478 16,523 SH   DFND 1 16,523 0 0
CACTUS INC A COMMON STOCK 127203107 65 2,276 SH   OTR 1 602 1,674 0
CADENCE DESIGN SYS INC COMMON STOCK 127387108 546 8,266 SH   DFND 1 7,695 0 571
CADENCE DESIGN SYS INC COMMON STOCK 127387108 50 769 SH   OTR 1 485 257 27
CADENCE BANCORP COMMON STOCK 12739A100 32 1,838 SH   DFND 1 469 0 1,369
CAESARS ENTERTAINMENT CORP COMMON STOCK 127686103 10 824 SH   DFND 1 662 0 162
CAL MAINE FOODS INC COMMON STOCK 128030202 466 11,662 SH   DFND 1 10,060 0 1,602
CAL MAINE FOODS INC COMMON STOCK 128030202 69 1,723 SH   OTR 1 1,723 0 0
CALAMOS CONV OPPORTUNITIES & I OTHER 128117108 36 3,481 SH   DFND 1 3,481 0 0
CALAMOS GLOBAL DYNAMIC INCOME OTHER 12811L107 2 319 SH   DFND 1 319 0 0
CALAMOS CONV & HIGH INCOME FD OTHER 12811P108 66 5,959 SH   OTR 1 5,959 0 0
CALAMOS STRATEGIC TOTAL RETURN OTHER 128125101 31 2,413 SH   DFND 1 2,413 0 0
CALAMP CORP COMMON STOCK 128126109 85 7,367 SH   DFND 1 5,955 0 1,412
CALAMP CORP COMMON STOCK 128126109 15 1,277 SH   OTR 1 1,277 0 0
CALAMP CORP COMMON STOCK 128126109 7 634 SH   DFND 5 634 0 0
CALAVO GROWERS INC COMMON STOCK 128246105 606 6,367 SH   DFND 1 6,367 0 0
CALAVO GROWERS INC COMMON STOCK 128246105 118 1,241 SH   OTR 1 686 555 0
CALERES INC COMMON STOCK 129500104 3 136 SH   DFND 1 136 0 0
CALERES INC COMMON STOCK 129500104 230 9,815 SH   OTR 1 9,815 0 0
CALIFORNIA RESOURCES CORP COMMON STOCK 13057Q206 1 145 SH   DFND 1 144 0 1
CALIFORNIA RESOURCES CORP COMMON STOCK 13057Q206 0 41 SH   OTR 1 33 0 8
CALIFORNIA WTR SVC GROUP COMMON STOCK 130788102 304 5,743 SH   DFND 1 5,743 0 0
CALIFORNIA WTR SVC GROUP COMMON STOCK 130788102 16 300 SH   OTR 1 300 0 0
CALLAWAY GOLF CO COMMON STOCK 131193104 2,479 127,740 SH   DFND 1 32,543 0 95,197
CALLAWAY GOLF CO COMMON STOCK 131193104 171 8,844 SH   OTR 1 6,347 2,497 0
CALLAWAY GOLF COMPANY COMMON STOCK 131193104 1 73 SH   DFND 5 73 0 0
CALLON PETE CO DEL COMMON STOCK 13123X102 237 54,410 SH   DFND 1 42,242 0 12,168
CALLON PETE CO DEL COMMON STOCK 13123X102 25 5,710 SH   OTR 1 4,719 991 0
CALLON PETROLEUM CO COMMON STOCK 13123X102 5 1,226 SH   DFND 5 1,226 0 0
CAMBREX CORP COMMON STOCK 132011107 387 6,499 SH   DFND 1 6,422 0 77
CAMBREX CORP COMMON STOCK 132011107 21 357 SH   OTR 1 357 0 0
CAMBRIA SHAREHOLDER YIELD ETF OTHER 132061201 11 300 SH   DFND 1 300 0 0
CAMBIA TRINITY OTHER 132061839 218 9,001 SH   DFND 5 0 0 9,001
CAMBRIDGE BANCORP COMMON STOCK 132152109 1,907 25,420 SH   DFND 1 25,420 0 0
CAMBRIDGE BANCORP COMMON STOCK 132152109 343 4,575 SH   OTR 1 4,575 0 0
CAMDEN NATL CORP COMMON STOCK 133034108 9 209 SH   DFND 1 181 0 28
CAMDEN PPTY TR SBI COMMON STOCK 133131102 665 5,988 SH   DFND 1 5,505 0 483
CAMDEN PPTY TR SBI COMMON STOCK 133131102 532 4,794 SH   OTR 1 4,544 5 245
CAMDEN PROPERTY TRUST COMMON STOCK 133131102 9 87 SH   DFND 5 87 0 0
CAMECO CORP COMMON STOCK 13321L108 128 13,485 SH   DFND 1 12,592 0 893
CAMECO CORP COMMON STOCK 13321L108 10 1,000 SH   OTR 1 0 0 1,000
CAMPBELL SOUP CO COMMON STOCK 134429109 1,001 21,325 SH   DFND 1 19,893 0 1,432
CAMPBELL SOUP CO COMMON STOCK 134429109 897 19,122 SH   OTR 1 18,994 29 99
CAMPING WORLD HOLDINGS INC A COMMON STOCK 13462K109 1 58 SH   DFND 1 58 0 0
CAMPING WORLD HOLDINGS INC A COMMON STOCK 13462K109 4 500 SH   OTR 1 500 0 0
CANADA GOOSE HOLDINGS INC COMMON STOCK 135086106 84 1,915 SH   DFND 1 1,915 0 0
CANADIAN IMPERIAL BK OF COMMER COMMON STOCK 136069101 436 5,281 SH   DFND 1 4,116 0 1,165
CANADIAN IMPERIAL BK OF COMMER COMMON STOCK 136069101 69 835 SH   OTR 1 726 20 89
CANADIAN IMPERIAL BK OF COMMRC COMMON STOCK 136069101 417 5,055 SH   DFND 5 5,055 0 0
CANADIAN NATL RY CO COMMON STOCK 136375102 11,372 126,550 SH   DFND 1 123,036 0 3,514
CANADIAN NATL RY CO COMMON STOCK 136375102 3,356 37,346 SH   OTR 1 35,434 701 1,211
CANADIAN NATIONAL RAILWAY CO COMMON STOCK 136375102 414 4,617 SH   DFND 5 4,617 0 0
CANADIAN NATURAL RESOURCES LTD COMMON STOCK 136385101 5,070 190,418 SH   DFND 1 186,720 0 3,698
CANADIAN NATURAL RESOURCES LTD COMMON STOCK 136385101 575 21,617 SH   OTR 1 21,223 0 394
CANADIAN NATURAL RESOURCES LTD COMMON STOCK 136385101 5 196 SH   DFND 5 196 0 0
CANADIAN PACIFIC RAILWAY LTD COMMON STOCK 13645T100 1,136 5,105 SH   DFND 1 4,866 0 239
CANADIAN PACIFIC RAILWAY LTD COMMON STOCK 13645T100 466 2,093 SH   OTR 1 1,575 246 272
CANADIAN PACIFIC RAILWAY LTD COMMON STOCK 13645T100 309 1,388 SH   DFND 5 1,388 0 0
CANADIAN SOLAR INC COMMON STOCK 136635109 1 75 SH   DFND 1 75 0 0
CANNAE HOLDINGS INC COMMON STOCK 13765N107 1 53 SH   DFND 1 53 0 0
CANNAE HOLDINGS INC COMMON STOCK 13765N107 9 325 SH   OTR 1 325 0 0
CANNTRUST HOLDINGS INC COMMON STOCK 137800207 1 1,145 SH   OTR 1 0 0 1,145
CANON INC SPONS A D R COMMON STOCK 138006309 1,176 44,051 SH   DFND 1 41,301 0 2,750
CANON INC SPONS A D R COMMON STOCK 138006309 17 637 SH   OTR 1 532 84 21
CANOPY GROWTH CORP COMMON STOCK 138035100 98 4,259 SH   DFND 1 4,259 0 0
CANOPY GROWTH CORP COMMON STOCK 138035100 39 1,689 SH   OTR 1 1,589 0 100
CANTEL MEDICAL CORP COMMON STOCK 138098108 746 9,958 SH   DFND 1 8,076 0 1,882
CANTEL MEDICAL CORP COMMON STOCK 138098108 418 5,583 SH   OTR 1 1,858 1,125 2,600
CANTEL MEDICAL CORPORATION COMMON STOCK 138098108 1 16 SH   DFND 5 16 0 0
CAPITAL ONE FINANCIAL CORP COMMON STOCK 14040H105 9,707 106,693 SH   DFND 1 102,342 0 4,351
CAPITAL ONE FINANCIAL CORP COMMON STOCK 14040H105 1,573 17,291 SH   OTR 1 11,321 877 5,093
CAPITAL ONE FINANCIAL CORP COMMON STOCK 14040H105 6 66 SH   DFND 5 66 0 0
CAPITOL FEDERAL FINANCIAL INC COMMON STOCK 14057J101 54 3,911 SH   DFND 1 3,911 0 0
CAPITOL FEDERAL FINANCIAL INC COMMON STOCK 14057J101 36 2,637 SH   OTR 1 2,637 0 0
CAPRICOR THERAPEUTICS INC COMMON STOCK 14070B309 28 7,462 SH   OTR 1 7,462 0 0
CARA THERAPEUTICS INC COMMON STOCK 140755109 4 200 SH   DFND 1 200 0 0
CARA THERAPEUTICS INC COMMON STOCK 140755109 18 1,000 SH   OTR 1 1,000 0 0
CARBO CERAMICS INC COMMON STOCK 140781105 2 889 SH   DFND 1 889 0 0
CARBON BLACK INC COMMON STOCK 14081R103 464 17,845 SH   DFND 1 17,625 0 220
CARBON BLACK INC COMMON STOCK 14081R103 268 10,330 SH   OTR 1 10,330 0 0
CARDINAL HEALTH INC COMMON STOCK 14149Y108 4,460 94,510 SH   DFND 1 93,685 0 825
CARDINAL HEALTH INC COMMON STOCK 14149Y108 1,113 23,580 SH   OTR 1 22,619 700 261
CARDINAL HEALTH INC COMMON STOCK 14149Y108 6 143 SH   DFND 5 143 0 0
CARDIOVASCULAR SYSTEMS INC COMMON STOCK 141619106 694 14,601 SH   DFND 1 14,601 0 0
CARDIOVASCULAR SYSTEMS INC COMMON STOCK 141619106 688 14,481 SH   OTR 1 13,783 698 0
CAREER ED CORP COMMON STOCK 141665109 91 5,698 SH   DFND 1 5,300 0 398
CAREDX INC COMMON STOCK 14167L103 12 528 SH   DFND 1 528 0 0
CARETRUST REIT INC COMMON STOCK 14174T107 8 320 SH   DFND 1 320 0 0
CARGURUS INC COMMON STOCK 141788109 0 12 SH   DFND 1 12 0 0
CARLISLE COS INC COMMON STOCK 142339100 1,086 7,462 SH   DFND 1 7,113 0 349
CARLISLE COS INC COMMON STOCK 142339100 124 853 SH   OTR 1 813 28 12
CARLISLE COMPANIES INC COMMON STOCK 142339100 23 159 SH   DFND 5 159 0 0
CARLYLE GROUP L P COMMON STOCK 14309L102 43 1,700 SH   OTR 1 1,700 0 0
CARMAX INC COMMON STOCK 143130102 2,962 33,659 SH   DFND 1 12,200 0 21,459
CARMAX INC COMMON STOCK 143130102 102 1,157 SH   OTR 1 22 1,094 41
CARMAX INC COMMON STOCK 143130102 221 2,517 SH   DFND 5 2,517 0 0
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CARNIVAL CORP COMMON STOCK 143658300 483 11,043 SH   OTR 1 9,175 1,739 129
CARNIVAL CORPORATION COMMON STOCK 143658300 5 116 SH   DFND 5 116 0 0
CARNIVAL PLC A D R COMMON STOCK 14365C103 362 8,622 SH   DFND 1 7,384 0 1,238
CARNIVAL PLC A D R COMMON STOCK 14365C103 61 1,437 SH   OTR 1 610 235 592
CAROLINA FINANCIAL CORP COMMON STOCK 143873107 133 3,742 SH   DFND 1 3,656 0 86
CAROLINA FINANCIAL CORP COMMON STOCK 143873107 21 601 SH   OTR 1 601 0 0
CAROLINA FINANCIAL CORP COMMON STOCK 143873107 10 291 SH   DFND 5 291 0 0
CARPENTER TECHNOLOGY CORP COMMON STOCK 144285103 154 2,985 SH   DFND 1 2,656 0 329
CARPENTER TECHNOLOGY CORP COMMON STOCK 144285103 61 1,188 SH   OTR 1 1,188 0 0
CARPENTER TECHNOLOGY CORP COMMON STOCK 144285103 10 211 SH   DFND 5 211 0 0
CARRIZO OIL & GAS INC COMMON STOCK 144577103 356 41,485 SH   DFND 1 35,774 0 5,711
CARRIZO OIL & GAS INC COMMON STOCK 144577103 76 8,830 SH   OTR 1 8,301 529 0
CARRIZO OIL & GAS INC COMMON STOCK 144577103 5 648 SH   DFND 5 648 0 0
CARS COM INC W I COMMON STOCK 14575E105 63 7,065 SH   DFND 1 5,892 0 1,173
CARS COM INC W I COMMON STOCK 14575E105 0 33 SH   OTR 1 33 0 0
CARTERS INC COMMON STOCK 146229109 640 7,016 SH   DFND 1 6,030 0 986
CARTERS INC COMMON STOCK 146229109 241 2,641 SH   OTR 1 1,447 71 1,123
CARTER INC COMMON STOCK 146229109 34 379 SH   DFND 5 379 0 0
CARVANA CO COMMON STOCK 146869102 33 500 SH   DFND 1 500 0 0
CARVANA CO COMMON STOCK 146869102 2 31 SH   OTR 1 0 0 31
CASELLA WASTE SYSTEMS INC A COMMON STOCK 147448104 148 3,448 SH   OTR 1 0 1,223 2,225
CASEYS GEN STORES INC COMMON STOCK 147528103 865 5,368 SH   DFND 1 5,343 0 25
CASEYS GEN STORES INC COMMON STOCK 147528103 25 155 SH   OTR 1 155 0 0
CASEYS GENERAL STORES INC COMMON STOCK 147528103 2 18 SH   DFND 5 18 0 0
CASS INFORMATION SYSTEMS INC COMMON STOCK 14808P109 228 4,210 SH   DFND 1 4,210 0 0
CASS INFORMATION SYSTEMS INC COMMON STOCK 14808P109 38 705 SH   OTR 1 705 0 0
CATALENT INC COMMON STOCK 148806102 27 557 SH   DFND 1 557 0 0
CATALENT INC COMMON STOCK 148806102 57 1,189 SH   OTR 1 0 1,152 37
CATALENT INC COMMON STOCK 148806102 59 1,245 SH   DFND 5 1,245 0 0
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CATERPILLAR INC COMMON STOCK 149123101 18,259 144,559 SH   OTR 1 133,484 1,095 9,980
CATERPILLAR INC COMMON STOCK 149123101 19 156 SH   DFND 5 106 0 50
CATCHMARK TIMBER TRUST INC A COMMON STOCK 14912Y202 22 2,105 SH   DFND 1 1,391 0 714
CATHAY GENERAL BANCORP COMMON STOCK 149150104 196 5,646 SH   DFND 1 5,646 0 0
CATO CORP CL A COMMON STOCK 149205106 14 814 SH   DFND 1 265 0 549
CATO CORP CL A COMMON STOCK 149205106 36 2,043 SH   OTR 1 2,000 43 0
CAVCO INDUSTRIES INC COMMON STOCK 149568107 57 298 SH   DFND 5 298 0 0
CEDAR FAIR LP COMMON STOCK 150185106 1,322 22,652 SH   DFND 1 21,352 0 1,300
CEDAR FAIR LP COMMON STOCK 150185106 92 1,570 SH   OTR 1 1,570 0 0
CELANESE CORP SER A COMMON STOCK 150870103 1,482 12,118 SH   DFND 1 11,663 0 455
CELANESE CORP SER A COMMON STOCK 150870103 502 4,111 SH   OTR 1 3,870 71 170
CELESTICA INC COMMON STOCK 15101Q108 21 2,874 SH   DFND 1 0 0 2,874
CELGENE CORP COMMON STOCK 151020104 65,520 659,826 SH   DFND 1 648,101 0 11,725
CELGENE CORP COMMON STOCK 151020104 14,670 147,741 SH   OTR 1 141,214 5,230 1,297
CELGENE CORP COMMON STOCK 151020104 2 26 SH   DFND 5 26 0 0
CELCUITY INC COMMON STOCK 15102K100 136 8,000 SH   DFND 1 8,000 0 0
CELCUITY INC COMMON STOCK 15102K100 601 35,417 SH   OTR 1 35,417 0 0
CELLECTAR BIOSCIENCES INC COMMON STOCK 15117F500 0 2 SH   DFND 1 2 0 0
CELLECTAR BIOSCIENCES INC COMMON STOCK 15117F500 6 2,857 SH   OTR 1 2,857 0 0
CELLECTIS A D R COMMON STOCK 15117K103 8 763 SH   DFND 1 461 0 302
CELLECTIS A D R COMMON STOCK 15117K103 11 1,061 SH   OTR 1 0 1,061 0
CELSIUS HOLDINGS INC COMMON STOCK 15118V207 52 15,000 SH   DFND 1 15,000 0 0
CEMEX SAB A D R COMMON STOCK 151290889 42 10,626 SH   DFND 1 10,626 0 0
CEMEX SAB A D R COMMON STOCK 151290889 23 5,940 SH   OTR 1 0 0 5,940
CENTENE CORP COMMON STOCK 15135B101 41,185 952,012 SH   DFND 1 945,579 0 6,433
CENTENE CORP COMMON STOCK 15135B101 5,929 137,074 SH   OTR 1 127,669 6,625 2,780
CENOVUS ENERGY INC COMMON STOCK 15135U109 633 67,492 SH   DFND 1 65,999 0 1,493
CENOVUS ENERGY INC COMMON STOCK 15135U109 37 3,918 SH   OTR 1 3,645 0 273
CENTENNIAL RESOURCE DEVELO A COMMON STOCK 15136A102 12 2,604 SH   DFND 1 1,759 0 845
CENTERPOINT ENERGY INC COMMON STOCK 15189T107 1,635 54,200 SH   DFND 1 50,932 0 3,268
CENTERPOINT ENERGY INC COMMON STOCK 15189T107 871 28,879 SH   OTR 1 28,320 559 0
CENTERPOINT ENERGY INC COMMON STOCK 15189T107 12 425 SH   DFND 5 425 0 0
CENTERSTATE BANK OF FLORIDA CO COMMON STOCK 15201P109 104 4,340 SH   DFND 1 3,725 0 615
CENTERSTATE BANK OF FLORIDA CO COMMON STOCK 15201P109 45 1,896 SH   OTR 1 89 1,807 0
CENTRAIS ELETRICAS BRASILEIRAS COMMON STOCK 15234Q207 12 1,254 SH   DFND 1 1,254 0 0
CENTRAL PACIFIC FINANCIAL CO COMMON STOCK 154760409 31 1,109 SH   DFND 1 1,109 0 0
CENTURYLINK INC COMMON STOCK 156700106 567 45,461 SH   DFND 1 39,817 0 5,644
CENTURYLINK INC COMMON STOCK 156700106 148 11,813 SH   OTR 1 7,648 229 3,936
CENTURYLINK INC COMMON STOCK 156700106 9 772 SH   DFND 5 772 0 0
CERNER CORPORATION COMMON STOCK 156782104 14,437 211,780 SH   DFND 1 211,290 0 490
CERNER CORPORATION COMMON STOCK 156782104 4,991 73,205 SH   OTR 1 72,321 515 369
CERNER CORP COMMON STOCK 156782104 26 382 SH   DFND 5 382 0 0
CERUS CORP COMMON STOCK 157085101 16 3,029 SH   OTR 1 0 0 3,029
CERUS CORP COMMON STOCK 157085101 1 265 SH   DFND 5 265 0 0
CEVA INC COMMON STOCK 157210105 49 1,658 SH   DFND 1 1,658 0 0
CHANGYOU COM LTD A D R COMMON STOCK 15911M107 170 17,871 SH   DFND 1 17,603 0 268
CHANGYOU.COM LTD COMMON STOCK 15911M107 1 123 SH   DFND 5 123 0 0
CHANNELADVISOR CORP COMMON STOCK 159179100 1 147 SH   DFND 1 147 0 0
CHARLES & COLVARD LTD COMMON STOCK 159765106 397 243,775 SH   DFND 1 41,500 0 202,275
CHARLES RIVER LABORATORIES COMMON STOCK 159864107 3,099 23,409 SH   DFND 1 23,270 0 139
CHARLES RIVER LABORATORIES COMMON STOCK 159864107 587 4,439 SH   OTR 1 4,217 181 41
CHARLES RIVER LABORATORIES COMMON STOCK 159864107 19 147 SH   DFND 5 147 0 0
CHART INDUSTRIES INC COMMON STOCK 16115Q308 92 1,482 SH   DFND 1 1,482 0 0
CHART INDUSTRIES INC COMMON STOCK 16115Q308 115 1,837 SH   OTR 1 1,057 780 0
CHARTER COMMUNICATIONS INC NEW COMMON STOCK 16119P108 2,521 6,119 SH   DFND 1 5,590 0 529
CHARTER COMMUNICATIONS INC NEW COMMON STOCK 16119P108 900 2,186 SH   OTR 1 1,497 381 308
CHASE CORP COMMON STOCK 16150R104 601 5,495 SH   DFND 1 5,495 0 0
CHASE CORP COMMON STOCK 16150R104 62 570 SH   OTR 1 570 0 0
CHATHAM LODGING TRUST COMMON STOCK 16208T102 5 248 SH   DFND 1 248 0 0
CHEESECAKE FACTORY INC COMMON STOCK 163072101 757 18,166 SH   DFND 1 18,166 0 0
CHEESECAKE FACTORY INC COMMON STOCK 163072101 161 3,851 SH   OTR 1 3,851 0 0
CHEFS WAREHOUSE HOLDINGS LLC COMMON STOCK 163086101 2,086 51,744 SH   DFND 1 51,744 0 0
CHEFS WAREHOUSE HOLDINGS LLC COMMON STOCK 163086101 291 7,218 SH   OTR 1 5,018 275 1,925
CHEGG INC COMMON STOCK 163092109 332 11,089 SH   DFND 1 11,089 0 0
CHEGG INC COMMON STOCK 163092109 49 1,630 SH   OTR 1 1,630 0 0
CHEMED CORP COMMON STOCK 16359R103 4,934 11,815 SH   DFND 1 11,693 0 122
CHEMED CORP COMMON STOCK 16359R103 574 1,375 SH   OTR 1 1,363 7 5
CHEMED CORPORATION COMMON STOCK 16359R103 34 82 SH   DFND 5 82 0 0
CHEMOURS COMPANY COMMON STOCK 163851108 107 7,104 SH   DFND 1 4,390 0 2,714
CHEMOURS COMPANY COMMON STOCK 163851108 55 3,695 SH   OTR 1 1,636 442 1,617
CHENIERE ENERGY PARTNERS LP COMMON STOCK 16411Q101 49 1,081 SH   DFND 1 0 0 1,081
CHENIERE ENERGY PARTNERS LP COMMON STOCK 16411Q101 45 1,000 SH   OTR 1 1,000 0 0
CHENIERE ENERGY INC COMMON STOCK 16411R208 1,251 19,830 SH   DFND 1 15,223 0 4,607
CHENIERE ENERGY INC COMMON STOCK 16411R208 106 1,695 SH   OTR 1 1,351 0 344
CHENIERE ENERGY INC COMMON STOCK 16411R208 6 100 SH   DFND 5 100 0 0
CHESAPEAKE ENERGY CORP COMMON STOCK 165167107 9 6,488 SH   DFND 1 5,193 0 1,295
CHESAPEAKE ENERGY CORP COMMON STOCK 165167107 0 60 SH   OTR 1 60 0 0
CHESAPEAKE UTILITIES CORP COMMON STOCK 165303108 240 2,521 SH   DFND 1 2,370 0 151
CHESAPEAKE UTILITIES CORP COMMON STOCK 165303108 4 39 SH   OTR 1 39 0 0
CHEVRON CORPORATION COMMON STOCK 166764100 166,609 1,404,797 SH   DFND 1 1,364,456 0 40,341
CHEVRON CORPORATION COMMON STOCK 166764100 95,476 805,030 SH   OTR 1 762,023 16,990 26,017
CHEVRON CORPORATION COMMON STOCK 166764100 1,115 9,406 SH   DFND 5 5,807 0 3,599
CHEWY INC CLASS A COMMON STOCK 16679L109 66 2,700 SH   DFND 1 2,700 0 0
CHIASMA INC COMMON STOCK 16706W102 5 1,000 SH   DFND 1 1,000 0 0
CHICOS FAS INC COMMON STOCK 168615102 4 1,045 SH   DFND 1 1,045 0 0
CHICOS FAS INC COMMON STOCK 168615102 0 73 SH   OTR 1 73 0 0
THE CHILDRENS PLACE, INC COMMON STOCK 168905107 30 394 SH   DFND 1 394 0 0
CHIMERA INVESTMENT CORP COMMON STOCK 16934Q208 42 2,171 SH   DFND 1 2,171 0 0
CHIMERA INVESTMENT CORP COMMON STOCK 16934Q208 143 7,289 SH   OTR 1 3,000 0 4,289
CHIMERA INVESTMENT CORP COMMON STOCK 16934Q208 112 5,742 SH   DFND 5 5,503 0 239
CHINA LIFE INSURANCE CO A D R COMMON STOCK 16939P106 1,260 109,368 SH   DFND 1 101,192 0 8,176
CHINA LIFE INSURANCE CO A D R COMMON STOCK 16939P106 101 8,744 SH   OTR 1 8,576 0 168
CHINA SOUTHN AIRLS LTD SPON A COMMON STOCK 169409109 15 499 SH   DFND 1 0 0 499
CHINA SOUTHN AIRLS LTD SPON A COMMON STOCK 169409109 5 164 SH   OTR 1 164 0 0
CHINA MOBILE LIMITED A D R COMMON STOCK 16941M109 3,395 82,014 SH   DFND 1 78,431 0 3,583
CHINA MOBILE LIMITED A D R COMMON STOCK 16941M109 706 17,081 SH   OTR 1 13,635 897 2,549
CHINA MOBILE LIMITED COMMON STOCK 16941M109 17 422 SH   DFND 5 422 0 0
CHINA PETROLEUM CHEM A D R COMMON STOCK 16941R108 94 1,594 SH   DFND 1 1,141 0 453
CHINA PETROLEUM CHEM A D R COMMON STOCK 16941R108 1 18 SH   OTR 1 0 0 18
CHINA TELECOM CORP LTD A D R COMMON STOCK 169426103 238 5,225 SH   DFND 1 4,588 0 637
CHINA TELECOM CORP LTD A D R COMMON STOCK 169426103 94 2,054 SH   OTR 1 0 114 1,940
CHINA TELECOM CORP LTD COMMON STOCK 169426103 15 337 SH   DFND 5 337 0 0
CHINA UNICOM HONG KONG LTD A D COMMON STOCK 16945R104 190 18,084 SH   DFND 1 14,477 0 3,607
CHINA UNICOM HONG KONG LTD A D COMMON STOCK 16945R104 5 444 SH   OTR 1 299 0 145
CHINA RAPID A D R COMMON STOCK 16953Q105 25 47,500 SH   DFND 1 47,500 0 0
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CHURCH AND DWIGHT CO INC COMMON STOCK 171340102 3,624 48,171 SH   OTR 1 42,788 320 5,063
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CIENA CORP COMMON STOCK 171779309 701 17,859 SH   DFND 1 16,165 0 1,694
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CRANE CO COMMON STOCK 224399105 703 8,715 SH   DFND 1 8,646 0 69
CRANE CO COMMON STOCK 224399105 102 1,259 SH   OTR 1 51 0 1,208
CREDIT SUISSE ASSET MGMT INCCO OTHER 224916106 3 1,000 SH   OTR 1 1,000 0 0
CREATIVE REALITIES INC COMMON STOCK 22530J200 0 4 SH   OTR 1 4 0 0
CREDIT ACCEPTANCE CORP COMMON STOCK 225310101 562 1,219 SH   DFND 1 1,219 0 0
CREDIT SUISSE GROUP A D R COMMON STOCK 225401108 1,156 94,697 SH   DFND 1 87,165 0 7,532
CREDIT SUISSE GROUP A D R COMMON STOCK 225401108 293 23,952 SH   OTR 1 19,280 1,674 2,998
CREDIT SUISSE GROUP AG COMMON STOCK 225401108 3 285 SH   DFND 5 285 0 0
CREE RESEARCH INC COMMON STOCK 225447101 18 365 SH   DFND 1 365 0 0
CREE RESEARCH INC COMMON STOCK 225447101 4 83 SH   OTR 1 83 0 0
CRESTWOOD EQUITY PARTNERS L P COMMON STOCK 226344208 41 1,135 SH   OTR 1 0 0 1,135
CRESTWOOD EQUITY PARTNERS LP COMMON STOCK 226344208 3 88 SH   DFND 5 88 0 0
CRITEO SA SPON A D R COMMON STOCK 226718104 3,402 181,993 SH   DFND 1 181,794 0 199
CRITEO SA SPON A D R COMMON STOCK 226718104 742 39,709 SH   OTR 1 39,709 0 0
CRITEO S A COMMON STOCK 226718104 37 2,004 SH   DFND 5 2,004 0 0
CROCS INC COMMON STOCK 227046109 301 10,841 SH   DFND 1 10,841 0 0
CROCS INC COMMON STOCK 227046109 623 22,452 SH   OTR 1 22,452 0 0
CRONOS GROUP INC COMMON STOCK 22717L101 20 2,250 SH   OTR 1 2,150 0 100
CROWDSTRIKE HOLDINGS INC A COMMON STOCK 22788C105 34 582 SH   DFND 1 582 0 0
CROWN CASTLE INTL CORP COMMON STOCK 22822V101 15,747 113,286 SH   DFND 1 107,182 0 6,104
CROWN CASTLE INTL CORP COMMON STOCK 22822V101 3,599 25,890 SH   OTR 1 23,088 561 2,241
CROWN CASTLE INTL CORP COMMON STOCK 22822V101 320 2,305 SH   DFND 5 2,305 0 0
CROWN CASTLE INTL 6.875 CV PR PREFERRED ST 22822V309 19 15 SH   DFND 1 15 0 0
CROWN HOLDINGS INC COMMON STOCK 228368106 142 2,157 SH   DFND 1 2,048 0 109
CROWN HOLDINGS INC COMMON STOCK 228368106 132 1,986 SH   OTR 1 1,487 246 253
CROWN HOLDINGS INC COMMON STOCK 228368106 5 78 SH   DFND 5 78 0 0
CRYOLIFE INC COMMON STOCK 228903100 2 56 SH   DFND 1 56 0 0
CRYOPORT INC COMMON STOCK 229050307 105 6,427 SH   DFND 1 6,427 0 0
CRYOPORT INC COMMON STOCK 229050307 156 9,530 SH   OTR 1 0 0 9,530
CTRIP COM INTERNATIONAL A D R COMMON STOCK 22943F100 520 17,763 SH   DFND 1 16,984 0 779
CTRIP COM INTERNATIONAL A D R COMMON STOCK 22943F100 38 1,283 SH   OTR 1 1,228 0 55
CTRIP.COM INTERNATIONAL COMMON STOCK 22943F100 2 90 SH   DFND 5 90 0 0
CUBESMART COMMON STOCK 229663109 495 14,186 SH   DFND 1 13,187 0 999
CUBESMART COMMON STOCK 229663109 207 5,924 SH   OTR 1 5,924 0 0
CUBESMART COMMON STOCK 229663109 8 250 SH   DFND 5 250 0 0
CUBIC CORP COMMON STOCK 229669106 471 6,687 SH   DFND 1 6,687 0 0
CUBIC CORP COMMON STOCK 229669106 87 1,239 SH   OTR 1 1,239 0 0
CULLEN FROST BANKERS INC COMMON STOCK 229899109 334 3,774 SH   DFND 1 3,381 0 393
CULLEN FROST BANKERS INC COMMON STOCK 229899109 66 744 SH   OTR 1 87 657 0
CULP INC COMMON STOCK 230215105 20 1,212 SH   DFND 1 1,212 0 0
CUMMINS INC COMMON STOCK 231021106 12,922 79,441 SH   DFND 1 78,016 0 1,425
CUMMINS INC COMMON STOCK 231021106 4,453 27,368 SH   OTR 1 20,990 1,370 5,008
CUMMINS INC COMMON STOCK 231021106 79 488 SH   DFND 5 488 0 0
CUMULUS MEDIA INC CL A COMMON STOCK 231082801 1 100 SH   DFND 1 100 0 0
CURTISS WRIGHT CORP COMMON STOCK 231561101 403 3,115 SH   DFND 1 3,073 0 42
CURTISS WRIGHT CORP COMMON STOCK 231561101 1,241 9,593 SH   OTR 1 7,861 0 1,732
CUSHING ROYALTY & INCOME FUND OTHER 23162T102 2 281 SH   DFND 5 281 0 0
CUSTOMERS BANCORP INC COMMON STOCK 23204G100 2 93 SH   DFND 1 93 0 0
CUSTOMERS BANCORP INC COMMON STOCK 23204G100 8 363 SH   OTR 1 0 363 0
CUTERA INC COMMON STOCK 232109108 29 1,000 SH   DFND 1 1,000 0 0
CYPRESS SEMICONDUCTOR CORP COMMON STOCK 232806109 150 6,447 SH   DFND 1 6,378 0 69
CYPRESS SEMICONDUCTOR CORP COMMON STOCK 232806109 407 17,436 SH   OTR 1 16,514 922 0
CYTOKINETICS INC COMMON STOCK 23282W605 2 190 SH   DFND 1 190 0 0
CYRUSONE INC COMMON STOCK 23283R100 1,339 16,927 SH   DFND 1 16,223 0 704
CYRUSONE INC COMMON STOCK 23283R100 216 2,725 SH   OTR 1 1,351 925 449
CYRUSONE INC COMMON STOCK 23283R100 7 91 SH   DFND 5 91 0 0
DMC GLOBAL INC COMMON STOCK 23291C103 44 1,011 SH   DFND 1 0 0 1,011
DEUTSCHE X-TRACKERS MSCI EM MK OTHER 233051101 17 785 SH   DFND 1 785 0 0
DEUTSCHE XTRACKERS MSCI EAFE E OTHER 233051200 8,210 253,873 SH   DFND 1 253,870 0 3
DEUTSCHE XTRACKERS MSCI EAFE E OTHER 233051200 3,431 106,105 SH   OTR 1 99,162 1,443 5,500
DBX MSCI EAFE CURRENCY HEDGED OTHER 233051200 6 187 SH   DFND 5 187 0 0
DEUTSCHE XTRACKERS USD HIGH ET OTHER 233051432 1 20 SH   DFND 1 20 0 0
DEUTSCHE X-TRACKERS HY CORP OTHER 233051432 221 4,426 SH   DFND 5 4,426 0 0
DEUTSCHE XTRACKERS MSCI JAPAN OTHER 233051507 1,275 32,029 SH   DFND 1 31,354 0 675
DEUTSCHE XTRACKERS MSCI JAPAN OTHER 233051507 603 15,141 SH   OTR 1 15,141 0 0
XTRACKERS MUNICIPAL INFRAS REV OTHER 233051705 15 525 SH   DFND 1 525 0 0
DEUTSCHE X TRACKERS MSCI EUROP OTHER 233051853 514 17,216 SH   DFND 1 17,216 0 0
DEUTSCHE X TRACKERS MSCI EUROP OTHER 233051853 1,043 34,956 SH   OTR 1 34,956 0 0
DBV TECHNOLOGIES SA A D R COMMON STOCK 23306J101 35 4,039 SH   OTR 1 0 952 3,087
DCP MIDSTREAM LP COMMON STOCK 23311P100 143 5,473 SH   DFND 1 4,209 0 1,264
DCP MIDSTREAM LP COMMON STOCK 23311P100 3 115 SH   OTR 1 115 0 0
DNP SELECT INCOME FUND INC OTHER 23325P104 1,178 91,570 SH   DFND 1 91,570 0 0
DNP SELECT INCOME FUND INC OTHER 23325P104 693 53,860 SH   OTR 1 50,160 0 3,700
DNP SELECT INCOME FUND INC OTHER 23325P104 41 3,190 SH   DFND 5 0 0 3,190
D R HORTON INC COMMON STOCK 23331A109 8,616 163,458 SH   DFND 1 158,610 0 4,848
D R HORTON INC COMMON STOCK 23331A109 1,186 22,498 SH   OTR 1 21,571 170 757
D.R. HORTON INC COMMON STOCK 23331A109 128 2,434 SH   DFND 5 2,434 0 0
D T E ENERGY CO COMMON STOCK 233331107 1,461 10,990 SH   DFND 1 10,273 0 717
D T E ENERGY CO COMMON STOCK 233331107 1,069 8,042 SH   OTR 1 7,397 631 14
DWS MUNICIPAL INCOME TRUST OTHER 233368109 40 3,489 SH   DFND 1 3,489 0 0
DWS MUNICIPAL INCOME TRUST OTHER 233368109 3 300 SH   OTR 1 300 0 0
DEUTSCHE STRATEGIC MUNICIPAL I OTHER 23342Q101 90 7,588 SH   DFND 1 7,588 0 0
DXC TECHNOLOGY CO COMMON STOCK 23355L106 854 28,968 SH   DFND 1 28,501 0 467
DXC TECHNOLOGY CO COMMON STOCK 23355L106 406 13,783 SH   OTR 1 13,372 0 411
DAKTRONICS INC COMMON STOCK 234264109 6 800 SH   DFND 1 800 0 0
DANA INCORPORATED COMMON STOCK 235825205 133 9,268 SH   DFND 1 9,268 0 0
DANA INCORPORATED COMMON STOCK 235825205 2 105 SH   OTR 1 0 0 105
DANAHER CORP COMMON STOCK 235851102 24,221 167,705 SH   DFND 1 162,443 0 5,262
DANAHER CORP COMMON STOCK 235851102 10,324 71,489 SH   OTR 1 55,805 2,228 13,456
DANAHER CORP COMMON STOCK 235851102 13 92 SH   DFND 5 92 0 0
DAQO NEW ENERGY CORP A D R COMMON STOCK 23703Q203 4 79 SH   DFND 1 79 0 0
DARDEN RESTAURANTS INC COMMON STOCK 237194105 3,979 33,656 SH   DFND 1 29,661 0 3,995
DARDEN RESTAURANTS INC COMMON STOCK 237194105 2,965 25,079 SH   OTR 1 24,857 82 140
DARDEN RESTAURANTS INC COMMON STOCK 237194105 26 225 SH   DFND 5 225 0 0
DARLING INGREDIENTS INC COMMON STOCK 237266101 468 24,450 SH   DFND 1 15,530 0 8,920
DASEKE INC COMMON STOCK 23753F107 1,547 618,974 SH   DFND 1 618,974 0 0
DATA I O CORP COMMON STOCK 237690102 35 9,015 SH   OTR 1 0 9,015 0
DAVE BUSTERS ENTERTAINMEN COMMON STOCK 238337109 401 10,320 SH   DFND 1 10,214 0 106
DAVE BUSTERS ENTERTAINMEN COMMON STOCK 238337109 63 1,610 SH   OTR 1 1,610 0 0
DAVITA INC COMMON STOCK 23918K108 160 2,813 SH   DFND 1 2,470 0 343
DAVITA INC COMMON STOCK 23918K108 32 572 SH   OTR 1 547 0 25
DAWSON GEOPHYSICAL CO COMMON STOCK 239360100 407 190,143 SH   DFND 1 0 0 190,143
DECIPHERA PHARMACEUTICALS IN COMMON STOCK 24344T101 22 648 SH   DFND 1 648 0 0
DECIPHERA PHARMACEUTICALS IN COMMON STOCK 24344T101 1 21 SH   OTR 1 21 0 0
DECKERS OUTDOOR CORP COMMON STOCK 243537107 30 202 SH   DFND 1 99 0 103
DECKERS OUTDOOR CORP COMMON STOCK 243537107 47 319 SH   OTR 1 0 182 137
DEERE CO COMMON STOCK 244199105 41,721 247,339 SH   DFND 1 243,868 0 3,471
DEERE CO COMMON STOCK 244199105 13,940 82,641 SH   OTR 1 77,176 507 4,958
DEERE & CO COMMON STOCK 244199105 28 168 SH   DFND 5 118 0 50
DELAWARE INVTS MINN MUN INCOME OTHER 24610V103 96 7,303 SH   DFND 1 7,303 0 0
DELAWARE INVTS MINN MUN INCOME OTHER 24610V103 1 107 SH   OTR 1 107 0 0
DELEK LOGISTICS PARTNERSHIP LP COMMON STOCK 24664T103 16 500 SH   OTR 1 500 0 0
DELEK HOLDCO INC COMMON STOCK 24665A103 455 12,566 SH   DFND 1 12,376 0 190
DELEK HOLDCO INC COMMON STOCK 24665A103 87 2,412 SH   OTR 1 2,028 0 384
DELL TECHNOLOGIES C COMMON STOCK 24703L202 390 7,512 SH   DFND 1 6,906 0 606
DELL TECHNOLOGIES C COMMON STOCK 24703L202 194 3,742 SH   OTR 1 1,499 239 2,004
DELTA AIR LINES INC COMMON STOCK 247361702 95,390 1,656,073 SH   DFND 1 1,632,169 0 23,904
DELTA AIR LINES INC COMMON STOCK 247361702 14,811 257,123 SH   OTR 1 244,717 8,739 3,667
DELTA AIR LINES COMMON STOCK 247361702 96 1,679 SH   DFND 5 1,679 0 0
DENBURY RESOURCES INC COMMON STOCK 247916208 7 5,478 SH   DFND 1 5,478 0 0
DELUXE CORP COMMON STOCK 248019101 2,812 57,193 SH   DFND 1 57,159 0 34
DELUXE CORP COMMON STOCK 248019101 412 8,365 SH   OTR 1 800 36 7,529
DENNYS CORP COMMON STOCK 24869P104 1,330 58,441 SH   DFND 1 52,548 0 5,893
DENNYS CORP COMMON STOCK 24869P104 280 12,287 SH   OTR 1 9,798 2,489 0
DENTSPLY SIRONA INC COMMON STOCK 24906P109 757 14,204 SH   DFND 1 13,377 0 827
DENTSPLY SIRONA INC COMMON STOCK 24906P109 193 3,610 SH   OTR 1 3,576 0 34
DENTSPLY SIRONA INC COMMON STOCK 24906P109 18 341 SH   DFND 5 341 0 0
DERMIRA INC COMMON STOCK 24983L104 1 133 SH   DFND 1 133 0 0
THE DESCARTES SYSTEMS GROUP IN COMMON STOCK 249906108 397 9,852 SH   DFND 1 6,616 0 3,236
THE DESCARTES SYSTEMS GROUP IN COMMON STOCK 249906108 303 7,508 SH   OTR 1 1,263 720 5,525
DESIGNER BRANDS INC COMMON STOCK 250565108 25 1,460 SH   OTR 1 0 1,460 0
DEVON ENERGY CORP COMMON STOCK 25179M103 732 30,389 SH   DFND 1 15,321 0 15,068
DEVON ENERGY CORP COMMON STOCK 25179M103 363 15,068 SH   OTR 1 4,185 0 10,883
DEXCOM INC COMMON STOCK 252131107 3,862 25,880 SH   DFND 1 25,803 0 77
DEXCOM INC COMMON STOCK 252131107 616 4,132 SH   OTR 1 1,123 2,982 27
DIAGEO PLC SPONSORED A D R COMMON STOCK 25243Q205 12,506 76,484 SH   DFND 1 69,785 0 6,699
DIAGEO PLC SPONSORED A D R COMMON STOCK 25243Q205 4,554 27,854 SH   OTR 1 25,998 961 895
DIAGEO PLC COMMON STOCK 25243Q205 35 219 SH   DFND 5 219 0 0
DIAMOND HILL INVT GROUP INC COMMON STOCK 25264R207 151 1,094 SH   DFND 1 1,094 0 0
DIAMOND OFFSHORE DRILLING INC COMMON STOCK 25271C102 4 756 SH   DFND 1 756 0 0
DIAMOND OFFSHORE DRILLING INC COMMON STOCK 25271C102 3 541 SH   OTR 1 0 541 0
DIAMONDROCK HOSPITALITY CO COMMON STOCK 252784301 12 1,217 SH   DFND 1 1,217 0 0
DIAMONDBACK ENERGY INC COMMON STOCK 25278X109 1,053 11,717 SH   DFND 1 11,302 0 415
DIAMONDBACK ENERGY INC COMMON STOCK 25278X109 524 5,828 SH   OTR 1 5,319 429 80
DIAMONDBACK ENERGY INC COMMON STOCK 25278X109 19 218 SH   DFND 5 218 0 0
DICKS SPORTING GOODS INC COMMON STOCK 253393102 760 18,631 SH   DFND 1 5,387 0 13,244
DICKS SPORTING GOODS INC COMMON STOCK 253393102 30 731 SH   OTR 1 650 31 50
DIEBOLD INCORPORATED COMMON STOCK 253651103 2 208 SH   DFND 1 208 0 0
DIGI INTERNATIONAL INC COMMON STOCK 253798102 16 1,194 SH   DFND 1 1,194 0 0
DIGITAL REALTY TRUST INC COMMON STOCK 253868103 27,541 212,163 SH   DFND 1 210,913 0 1,250
DIGITAL REALTY TRUST INC COMMON STOCK 253868103 2,884 22,211 SH   OTR 1 21,663 31 517
DIGITAL REALTY TRUST INC COMMON STOCK 253868103 38 296 SH   DFND 5 296 0 0
DIME CMNTY BANCORP INC COMMON STOCK 253922108 4 207 SH   DFND 1 207 0 0
DILLARDS INC CL A COMMON STOCK 254067101 568 8,594 SH   DFND 1 232 0 8,362
DINE BRANDS GLOBAL INC COMMON STOCK 254423106 88 1,156 SH   DFND 1 1,119 0 37
DINE BRANDS GLOBAL INC COMMON STOCK 254423106 3 46 SH   OTR 1 0 46 0
DIODES INC COMMON STOCK 254543101 3,300 82,186 SH   DFND 1 80,564 0 1,622
DIODES INC COMMON STOCK 254543101 581 14,484 SH   OTR 1 14,484 0 0
DIODES INC COMMON STOCK 254543101 16 417 SH   DFND 5 417 0 0
DIREXION DAILY 20 YEAR TRSY BE OTHER 25459Y678 12 1,150 SH   DFND 1 1,150 0 0
DIREXION ALL CAP INSIDER ETF OTHER 25459Y769 21 550 SH   DFND 1 550 0 0
DIREXION SHS ETF TR ETF OTHER 25460E133 1 30 SH   OTR 1 30 0 0
WALT DISNEY CO THE COMMON STOCK 254687106 127,694 979,855 SH   DFND 1 917,107 0 62,748
WALT DISNEY CO THE COMMON STOCK 254687106 48,928 375,449 SH   OTR 1 340,951 11,200 23,298
WALT DISNEY CO COMMON STOCK 254687106 528 4,056 SH   DFND 5 3,631 0 425
DISCOVER FINL SVCS COMMON STOCK 254709108 4,459 54,978 SH   DFND 1 52,144 0 2,834
DISCOVER FINL SVCS COMMON STOCK 254709108 2,839 35,007 SH   OTR 1 29,613 1,485 3,909
DISCOVER FINANCIAL SERVICES COMMON STOCK 254709108 62 768 SH   DFND 5 535 0 233
DISCOVERY INC A COMMON STOCK 25470F104 1,972 74,062 SH   DFND 1 73,566 0 496
DISCOVERY INC A COMMON STOCK 25470F104 27 1,023 SH   OTR 1 1,023 0 0
DISCOVERY COMMUNICATIONS COMMON STOCK 25470F104 40 1,536 SH   DFND 5 1,536 0 0
DISCOVERY INC C COMMON STOCK 25470F302 565 22,949 SH   DFND 1 22,949 0 0
DISCOVERY INC C COMMON STOCK 25470F302 881 35,810 SH   OTR 1 7,120 0 28,690
DISH NETWORK CORP CL A COMMON STOCK 25470M109 100 2,947 SH   DFND 1 2,261 0 686
DISH NETWORK CORP CL A COMMON STOCK 25470M109 3 93 SH   OTR 1 0 84 9
DISH NETWORK CORP COMMON STOCK 25470M109 1 54 SH   DFND 5 54 0 0
DR REDDYS LABORATORIES LTD A D COMMON STOCK 256135203 321 8,461 SH   DFND 1 6,444 0 2,017
DR REDDYS LABORATORIES LTD A D COMMON STOCK 256135203 26 684 SH   OTR 1 291 75 318
DOCUSIGN INC COMMON STOCK 256163106 2,513 40,587 SH   DFND 1 40,587 0 0
DOCUSIGN INC COMMON STOCK 256163106 230 3,704 SH   OTR 1 3,578 126 0
DOLBY LABORATORIES INC CL A COMMON STOCK 25659T107 163 2,528 SH   DFND 1 2,324 0 204
DOLBY LABORATORIES INC CL A COMMON STOCK 25659T107 163 2,516 SH   OTR 1 101 0 2,415
DOLBY LABORATORIES INC COMMON STOCK 25659T107 15 247 SH   DFND 5 247 0 0
DOLLAR GENERAL CORP COMMON STOCK 256677105 3,864 24,311 SH   DFND 1 23,005 0 1,306
DOLLAR GENERAL CORP COMMON STOCK 256677105 1,561 9,815 SH   OTR 1 7,746 1,965 104
DOLLAR GENERAL CORP COMMON STOCK 256677105 197 1,239 SH   DFND 5 1,239 0 0
DOLLAR TREE INC COMMON STOCK 256746108 16,285 142,650 SH   DFND 1 138,923 0 3,727
DOLLAR TREE INC COMMON STOCK 256746108 5,450 47,739 SH   OTR 1 47,114 128 497
DOLLAR TREE INC COMMON STOCK 256746108 11 104 SH   DFND 5 104 0 0
DOMINION ENERGY INC COMMON STOCK 25746U109 10,839 133,747 SH   DFND 1 126,194 0 7,553
DOMINION ENERGY INC COMMON STOCK 25746U109 5,082 62,702 SH   OTR 1 62,488 130 84
DOMINION RESOURCES INC/VA COMMON STOCK 25746U109 447 5,516 SH   DFND 5 5,516 0 0
DOMINOS PIZZA INC COMMON STOCK 25754A201 601 2,458 SH   DFND 1 933 0 1,525
DOMINOS PIZZA INC COMMON STOCK 25754A201 151 617 SH   OTR 1 359 207 51
DOMTAR CORP COMMON STOCK 257559203 29 814 SH   DFND 1 483 0 331
DONALDSON CO INC COMMON STOCK 257651109 26,900 516,515 SH   DFND 1 490,422 0 26,093
DONALDSON CO INC COMMON STOCK 257651109 18,821 361,379 SH   OTR 1 175,081 8,096 178,202
RR DONNELLEY SONS CO COMMON STOCK 257867200 1 385 SH   DFND 1 385 0 0
RR DONNELLEY SONS CO COMMON STOCK 257867200 0 106 SH   OTR 1 106 0 0
DONNELLEY FINANCIAL SOLUTIONS COMMON STOCK 25787G100 2 144 SH   DFND 1 144 0 0
DORCHESTER MINERALS L P COMMON STOCK 25820R105 45 2,400 SH   OTR 1 2,400 0 0
DORMAN PRODUCTS INC COMMON STOCK 258278100 1,266 15,921 SH   DFND 1 14,503 0 1,418
DORMAN PRODUCTS INC COMMON STOCK 258278100 210 2,644 SH   OTR 1 1,219 175 1,250
DORMAN PRODUCTS INC COMMON STOCK 258278100 72 908 SH   DFND 5 908 0 0
DOUGLAS EMMITT INC COMMON STOCK 25960P109 28 657 SH   DFND 1 575 0 82
DOUGLAS EMMETT INC COMMON STOCK 25960P109 1 29 SH   DFND 5 29 0 0
DOUGLAS DYNAMICS INC COMMON STOCK 25960R105 102 2,286 SH   DFND 1 2,250 0 36
DOUGLAS DYNAMICS INC COMMON STOCK 25960R105 149 3,340 SH   OTR 1 0 390 2,950
DOVER CORP COMMON STOCK 260003108 11,780 118,331 SH   DFND 1 116,507 0 1,824
DOVER CORP COMMON STOCK 260003108 4,220 42,393 SH   OTR 1 41,926 200 267
DOVER CORP COMMON STOCK 260003108 37 379 SH   DFND 5 379 0 0
DOW INC COMMON STOCK 260557103 17,138 359,663 SH   DFND 1 353,794 0 5,869
DOW INC COMMON STOCK 260557103 11,061 232,137 SH   OTR 1 222,258 3,173 6,706
DOW INC COMMON STOCK 260557103 42 893 SH   DFND 5 893 0 0
DRIL QUIP INC COMMON STOCK 262037104 435 8,658 SH   DFND 1 8,658 0 0
DRIL QUIP INC COMMON STOCK 262037104 74 1,478 SH   OTR 1 1,478 0 0
DRIL-QUIP COMMON STOCK 262037104 16 326 SH   DFND 5 326 0 0
DRIVE SHACK INC COMMON STOCK 262077100 17 4,000 SH   DFND 1 4,000 0 0
DROPBOX INC CLASS A COMMON STOCK 26210C104 205 10,168 SH   DFND 1 8,168 0 2,000
DROPBOX INC CLASS A COMMON STOCK 26210C104 11 525 SH   OTR 1 525 0 0
DUFF & PHELPS UTIL & CORPORATE OTHER 26432K108 13 1,400 SH   OTR 1 1,400 0 0
DUFF PHELPS SELECT MLP AND MID COMMON STOCK 26433F108 3 750 SH   DFND 1 750 0 0
DUKE REALTY CORPORATION COMMON STOCK 264411505 279 8,224 SH   DFND 1 7,623 0 601
DUKE REALTY CORPORATION COMMON STOCK 264411505 49 1,432 SH   OTR 1 1,353 36 43
DUKE REALTY CORP COMMON STOCK 264411505 6 198 SH   DFND 5 198 0 0
DUKE ENERGY HOLDING CORP COMMON STOCK 26441C204 24,468 255,246 SH   DFND 1 243,760 0 11,486
DUKE ENERGY HOLDING CORP COMMON STOCK 26441C204 11,361 118,519 SH   OTR 1 109,352 1,427 7,740
DUKE ENERGY CORP COMMON STOCK 26441C204 460 4,802 SH   DFND 5 4,802 0 0
DULUTH HOLDINGS INC CL B COMMON STOCK 26443V101 171 20,183 SH   DFND 1 20,183 0 0
DULUTH HOLDINGS INC CL B COMMON STOCK 26443V101 47 5,564 SH   OTR 1 5,564 0 0
DUNKIN BRANDS GROUP INC COMMON STOCK 265504100 439 5,528 SH   DFND 1 578 0 4,950
DUNKIN BRANDS GROUP INC COMMON STOCK 265504100 213 2,694 SH   OTR 1 950 67 1,677
DUPONT DE NEMOURS INC WI COMMON STOCK 26614N102 20,205 283,344 SH   DFND 1 277,265 0 6,079
DUPONT DE NEMOURS INC WI COMMON STOCK 26614N102 16,125 226,126 SH   OTR 1 216,771 3,688 5,667
DUPONT DE NEMOURS INC COMMON STOCK 26614N102 52 732 SH   DFND 5 732 0 0
DYCOM INDS INC COMMON STOCK 267475101 748 14,647 SH   DFND 1 14,406 0 241
DYCOM INDS INC COMMON STOCK 267475101 126 2,465 SH   OTR 1 2,465 0 0
DYNEX CAPITAL INC COMMON STOCK 26817Q886 51 3,462 SH   DFND 1 243 0 3,219
ECA MARCELLUS TR I COMMON STOCK 26827L109 6 5,000 SH   DFND 1 5,000 0 0
ECA MARCELLUS TR I COMMON STOCK 26827L109 6 5,000 SH   OTR 1 5,000 0 0
EDAP TMS SA A D R COMMON STOCK 268311107 9 1,925 SH   DFND 1 1,925 0 0
E L F BEAUTY INC COMMON STOCK 26856L103 1 76 SH   DFND 1 76 0 0
EL POLLO LOCO HOLDINGS INC COMMON STOCK 268603107 2 161 SH   DFND 1 161 0 0
E N I SPA A D R COMMON STOCK 26874R108 388 12,725 SH   DFND 1 8,233 0 4,492
E N I SPA A D R COMMON STOCK 26874R108 45 1,457 SH   OTR 1 122 207 1,128
ENI SPA COMMON STOCK 26874R108 2 87 SH   DFND 5 87 0 0
E O G RES INC COMMON STOCK 26875P101 6,433 86,673 SH   DFND 1 80,483 0 6,190
E O G RES INC COMMON STOCK 26875P101 1,161 15,638 SH   OTR 1 10,844 2,975 1,819
EOG RESOURCES INC COMMON STOCK 26875P101 6 93 SH   DFND 5 93 0 0
EQT CORP COMMON STOCK 26884L109 52 4,846 SH   DFND 1 4,135 0 711
EQT CORP COMMON STOCK 26884L109 21 2,000 SH   OTR 1 0 2,000 0
EPR PROPERTIES COMMON STOCK 26884U109 736 9,583 SH   DFND 1 9,231 0 352
EPR PROPERTIES COMMON STOCK 26884U109 131 1,700 SH   OTR 1 1,641 0 59
EPR PROPERTIES COMMON STOCK 26884U109 12 163 SH   DFND 5 163 0 0
EQM MIDSTREAM PARTNERS LP COMMON STOCK 26885B100 447 13,657 SH   DFND 1 1,494 0 12,163
EQT MIDSTREAM PARTNERS LP COMMON STOCK 26885B100 2 79 SH   DFND 5 79 0 0
EAGLE BANCORP INC COMMON STOCK 268948106 90 2,016 SH   DFND 1 823 0 1,193
US VEGAN CLIMATE ETF OTHER 26922A297 1 40 SH   DFND 1 40 0 0
CBOE VESTS P500 DIV ARSTCRTS T OTHER 26922A537 143 3,250 SH   DFND 1 3,250 0 0
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EBIX INC COMMON STOCK 278715206 129 3,062 SH   DFND 1 3,062 0 0
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ECHOSTAR CORP A COMMON STOCK 278768106 27 673 SH   DFND 1 673 0 0
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ECOLAB INC COMMON STOCK 278865100 57,304 289,358 SH   OTR 1 245,469 4,845 39,044
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ECOPETROL SA A D R COMMON STOCK 279158109 3 173 SH   OTR 1 173 0 0
EDGEWELL PERSONAL CARE CO COMMON STOCK 28035Q102 854 26,271 SH   DFND 1 24,819 0 1,452
EDGEWELL PERSONAL CARE CO COMMON STOCK 28035Q102 93 2,872 SH   OTR 1 2,872 0 0
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EDISON INTERNATIONAL COMMON STOCK 281020107 1,776 23,542 SH   OTR 1 21,499 2,043 0
EDISON INTERNATIONAL COMMON STOCK 281020107 20 273 SH   DFND 5 273 0 0
EDITAS MEDICINE INC COMMON STOCK 28106W103 36 1,595 SH   DFND 1 1,595 0 0
EDITAS MEDICINE INC COMMON STOCK 28106W103 3 150 SH   OTR 1 150 0 0
EDWARDS LIFESCIENCES CORP COMMON STOCK 28176E108 42,885 195,010 SH   DFND 1 191,714 0 3,296
EDWARDS LIFESCIENCES CORP COMMON STOCK 28176E108 14,396 65,463 SH   OTR 1 52,363 5,123 7,977
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EHEALTH INC COMMON STOCK 28238P109 1 14 SH   OTR 1 14 0 0
8X8 INC COMMON STOCK 282914100 5 258 SH   DFND 1 258 0 0
EL PASO ELEC CO COMMON STOCK 283677854 794 11,837 SH   DFND 1 11,533 0 304
EL PASO ELEC CO COMMON STOCK 283677854 100 1,491 SH   OTR 1 1,483 8 0
EL PASO ENERGY CAP TR PFD CV 4 PREFERRED ST 283678209 26 500 SH   DFND 1 0 0 500
ELANCO ANIMAL HEALTH INC COMMON STOCK 28414H103 312 11,720 SH   DFND 1 11,536 0 184
ELANCO ANIMAL HEALTH INC COMMON STOCK 28414H103 462 17,378 SH   OTR 1 3,868 0 13,510
ELDORADO GOLD CORP COMMON STOCK 284902509 1,493 192,160 SH   DFND 1 191,201 0 959
ELECTRONIC ARTS INC COMMON STOCK 285512109 31,761 324,688 SH   DFND 1 322,771 0 1,917
ELECTRONIC ARTS INC COMMON STOCK 285512109 3,774 38,585 SH   OTR 1 34,510 4,019 56
ELECTRONIC ARTS INC COMMON STOCK 285512109 11 117 SH   DFND 5 117 0 0
ELEMENT SOLUTIONS INC COMMON STOCK 28618M106 5 508 SH   DFND 1 508 0 0
EMBOTELLADORA ANDINA SPON A D COMMON STOCK 29081P204 10 551 SH   DFND 1 551 0 0
EMBOTELLADORA ANDINA B A D R COMMON STOCK 29081P303 23 1,137 SH   DFND 1 991 0 146
EMBOTELLADORA ANDINA B A D R COMMON STOCK 29081P303 5 272 SH   OTR 1 272 0 0
EMBOTELLADORA ANDINA SA COMMON STOCK 29081P303 3 170 SH   DFND 5 170 0 0
EMBRAER SA A D R COMMON STOCK 29082A107 113 6,525 SH   DFND 1 5,337 0 1,188
EMBRAER SA A D R COMMON STOCK 29082A107 26 1,535 SH   OTR 1 125 0 1,410
EMCORE CORPORATION COMMON STOCK 290846203 0 150 SH   DFND 1 150 0 0
EMCOR GROUP INC COMMON STOCK 29084Q100 1,716 19,917 SH   DFND 1 18,818 0 1,099
EMCOR GROUP INC COMMON STOCK 29084Q100 338 3,922 SH   OTR 1 1,843 0 2,079
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EMERGENT BIOSOLUTIONS INC COMMON STOCK 29089Q105 292 5,578 SH   DFND 1 5,411 0 167
EMERGENT BIOSOLUTIONS INC COMMON STOCK 29089Q105 117 2,246 SH   OTR 1 189 2,057 0
EMERSON ELECTRIC CO COMMON STOCK 291011104 95,999 1,435,817 SH   DFND 1 1,402,242 0 33,575
EMERSON ELECTRIC CO COMMON STOCK 291011104 87,031 1,301,688 SH   OTR 1 1,223,806 21,191 56,691
EMERSON ELECTRIC CO COMMON STOCK 291011104 14 214 SH   DFND 5 214 0 0
EMERALD EXPOSITIONS EVENTS I COMMON STOCK 29103B100 6 575 SH   DFND 1 575 0 0
EMERSON RADIO CORP COMMON STOCK 291087203 2 2,000 SH   DFND 1 2,000 0 0
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EMPIRE STATE REALTY TRUST IN COMMON STOCK 292104106 275 19,276 SH   DFND 1 19,276 0 0
EMPIRE STATE REALTY TRUST IN COMMON STOCK 292104106 57 4,016 SH   OTR 1 4,016 0 0
EMPLOYERS HOLDINGS INC COMMON STOCK 292218104 12 271 SH   DFND 1 207 0 64
ENCANA CORP COMMON STOCK 292505104 520 113,018 SH   DFND 1 99,245 0 13,773
ENCANA CORP COMMON STOCK 292505104 96 20,826 SH   OTR 1 20,438 0 388
ENBRIDGE INC COMMON STOCK 29250N105 24,919 710,349 SH   DFND 1 690,166 0 20,183
ENBRIDGE INC COMMON STOCK 29250N105 8,153 232,391 SH   OTR 1 201,934 922 29,535
ENBRIDGE INC COMMON STOCK 29250N105 553 15,792 SH   DFND 5 15,792 0 0
ENANTA PHARMACEUTICALS INC COMMON STOCK 29251M106 13 222 SH   DFND 1 222 0 0
ENCORE CAPITAL GROUP INC COMMON STOCK 292554102 8 226 SH   DFND 1 226 0 0
ENCORE CAPITAL GROUP INC COMMON STOCK 292554102 2 59 SH   OTR 1 0 59 0
ENCOMPASS HEALTH CORPORATION COMMON STOCK 29261A100 254 4,009 SH   DFND 1 3,763 0 246
ENCOMPASS HEALTH CORPORATION COMMON STOCK 29261A100 190 3,003 SH   OTR 1 2,030 277 696
ENCOMPASS HEALTH CORP COMMON STOCK 29261A100 39 628 SH   DFND 5 628 0 0
ENERGY FUELS INC COMMON STOCK 292671708 0 35 SH   DFND 1 35 0 0
ENERGY RECOVERY INC COMMON STOCK 29270J100 6 700 SH   OTR 1 700 0 0
ENERGIZER HLDGS INC COMMON STOCK 29272W109 2,046 46,947 SH   DFND 1 45,684 0 1,263
ENERGIZER HLDGS INC COMMON STOCK 29272W109 689 15,796 SH   OTR 1 14,410 1,086 300
ENERGIZER HOLDINGS INC COMMON STOCK 29272W109 1 23 SH   DFND 5 23 0 0
ENERGY TRANSFER EQUITY L P COMMON STOCK 29273V100 2,432 185,931 SH   DFND 1 109,346 0 76,585
ENERGY TRANSFER EQUITY L P COMMON STOCK 29273V100 971 74,246 SH   OTR 1 50,143 5,254 18,849
ENERGY TRANSFER EQUITY LP COMMON STOCK 29273V100 5 442 SH   DFND 5 442 0 0
ENEL AMERICAS SA A D R COMMON STOCK 29274F104 1,436 157,651 SH   DFND 1 155,415 0 2,236
ENEL AMERICAS SA A D R COMMON STOCK 29274F104 267 29,372 SH   OTR 1 29,028 19 325
ENERSIS S.A. COMMON STOCK 29274F104 12 1,367 SH   DFND 5 1,367 0 0
ENERSYS COMMON STOCK 29275Y102 774 11,728 SH   DFND 1 11,470 0 258
ENERSYS COMMON STOCK 29275Y102 114 1,723 SH   OTR 1 1,667 0 56
ENERPLUS CORP COMMON STOCK 292766102 244 32,743 SH   DFND 1 2,282 0 30,461
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ENEL CHILE SA A D R COMMON STOCK 29278D105 21 4,981 SH   OTR 1 859 731 3,391
ENLINK MIDSTREAM LLC COMMON STOCK 29336T100 2 256 SH   DFND 5 256 0 0
ENNIS INC COMMON STOCK 293389102 59 2,909 SH   DFND 1 2,909 0 0
ENNIS INC COMMON STOCK 293389102 40 2,000 SH   OTR 1 2,000 0 0
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ENPHASE ENERGY INC COMMON STOCK 29355A107 1 27 SH   OTR 1 27 0 0
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ENSIGN GROUP INC THE COMMON STOCK 29358P101 521 10,991 SH   DFND 1 10,991 0 0
ENSIGN GROUP INC THE COMMON STOCK 29358P101 38 799 SH   OTR 1 799 0 0
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ENTEGRIS INC COMMON STOCK 29362U104 415 8,812 SH   OTR 1 3,173 1,368 4,271
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ENTERCOM COMMUNICATIONS CORP C COMMON STOCK 293639100 13 3,947 SH   DFND 1 3,806 0 141
ENTERGY CORPORATION COMMON STOCK 29364G103 877 7,473 SH   DFND 1 6,041 0 1,432
ENTERGY CORPORATION COMMON STOCK 29364G103 523 4,451 SH   OTR 1 4,368 61 22
ENTERPRISE FINL SVCS COMMON STOCK 293712105 163 3,997 SH   OTR 1 0 965 3,032
ENTERPRISE PRODUCTS PARTNERS COMMON STOCK 293792107 4,460 156,056 SH   DFND 1 104,297 0 51,759
ENTERPRISE PRODUCTS PARTNERS COMMON STOCK 293792107 2,676 93,638 SH   OTR 1 70,949 11,600 11,089
ENTERPRISE PRODUCTS PARTNERS COMMON STOCK 293792107 95 3,336 SH   DFND 5 3,336 0 0
ENTREE RESOURCES LTD COMMON STOCK 29384J103 0 500 SH   DFND 1 500 0 0
ENVESTNET INC COMMON STOCK 29404K106 186 3,285 SH   DFND 1 802 0 2,483
ENVESTNET INC COMMON STOCK 29404K106 653 11,513 SH   OTR 1 0 9,845 1,668
ENVESTNET INC COMMON STOCK 29404K106 31 551 SH   DFND 5 551 0 0
EPAM SYSTEMS INC COMMON STOCK 29414B104 33 181 SH   DFND 1 81 0 100
EPAM SYSTEMS INC COMMON STOCK 29414B104 476 2,608 SH   OTR 1 1,258 476 874
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E PLUS INC COMMON STOCK 294268107 105 1,383 SH   DFND 1 23 0 1,360
E PLUS INC COMMON STOCK 294268107 56 731 SH   OTR 1 0 688 43
EQUIFAX INC COMMON STOCK 294429105 1,833 13,033 SH   DFND 1 12,237 0 796
EQUIFAX INC COMMON STOCK 294429105 239 1,699 SH   OTR 1 1,558 128 13
EQUIFAX INCORPORATED COMMON STOCK 294429105 3 27 SH   DFND 5 27 0 0
EQUINIX INC COMMON STOCK 29444U700 3,390 5,878 SH   DFND 1 5,635 0 243
EQUINIX INC COMMON STOCK 29444U700 1,152 1,997 SH   OTR 1 1,395 455 147
EQUINIX INC COMMON STOCK 29444U700 266 461 SH   DFND 5 461 0 0
EQUINOR ASA SPON ADR COMMON STOCK 29446M102 655 34,563 SH   DFND 1 27,766 0 6,797
EQUINOR ASA SPON ADR COMMON STOCK 29446M102 111 5,863 SH   OTR 1 2,455 2,311 1,097
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EQUITRANS MIDSTREAM CORP COMMON STOCK 294600101 17 1,200 SH   OTR 1 0 1,200 0
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EQUITY COMMONWEALTH COMMON STOCK 294628102 127 3,701 SH   DFND 1 2,948 0 753
EQUITY COMMONWEALTH COMMON STOCK 294628102 155 4,539 SH   OTR 1 0 110 4,429
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EQUITY LIFESTYLE PPTYS INC COMMON STOCK 29472R108 229 1,715 SH   OTR 1 1,637 63 15
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EQUITY RESIDENTIAL COMMON STOCK 29476L107 4,459 51,691 SH   DFND 1 50,290 0 1,401
EQUITY RESIDENTIAL COMMON STOCK 29476L107 1,332 15,441 SH   OTR 1 12,636 0 2,805
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ERICCSON (LM) TEL COMMON STOCK 294821608 4 544 SH   DFND 5 544 0 0
ERIE INDEMNITY CO COMMON STOCK 29530P102 133 714 SH   DFND 1 667 0 47
ERIE INDEMNITY CO COMMON STOCK 29530P102 21 112 SH   OTR 1 0 56 56
E S C O TECHNOLOGIES INC COMMON STOCK 296315104 1,764 22,170 SH   DFND 1 20,677 0 1,493
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ESSEX PROPERTY TRUST INC COMMON STOCK 297178105 242 740 SH   OTR 1 727 3 10
ESSEX PROPERTY TRUST INC COMMON STOCK 297178105 20 64 SH   DFND 5 64 0 0
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ETSY INC COMMON STOCK 29786A106 2,345 41,498 SH   DFND 1 41,311 0 187
ETSY INC COMMON STOCK 29786A106 544 9,629 SH   OTR 1 7,587 32 2,010
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EURONET WORLDWIDE INC COMMON STOCK 298736109 866 5,922 SH   DFND 1 5,793 0 129
EURONET WORLDWIDE INC COMMON STOCK 298736109 638 4,360 SH   OTR 1 3,579 563 218
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EVERCORE INC COMMON STOCK 29977A105 1,193 14,892 SH   DFND 1 14,573 0 319
EVERCORE INC COMMON STOCK 29977A105 377 4,706 SH   OTR 1 3,887 0 819
EVERCORE PARTNERS INC COMMON STOCK 29977A105 1 13 SH   DFND 5 13 0 0
EVERBRIDGE INC COMMON STOCK 29978A104 283 4,580 SH   DFND 1 4,580 0 0
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EVERGY INC COMMON STOCK 30034W106 2,034 30,558 SH   DFND 1 28,461 0 2,097
EVERGY INC COMMON STOCK 30034W106 1,252 18,813 SH   OTR 1 16,443 75 2,295
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EVERTEC INC COMMON STOCK 30040P103 50 1,608 SH   OTR 1 0 1,608 0
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EVERSOURCE ENERGY COMMON STOCK 30040W108 1,492 17,461 SH   OTR 1 14,208 1,329 1,924
EVERSOURCE ENERGY COMMON STOCK 30040W108 39 465 SH   DFND 5 465 0 0
EVERSPIN TECHNOLOGIES INC COMMON STOCK 30041T104 2 300 SH   DFND 1 300 0 0
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EVOLENT HEALTH INC A COMMON STOCK 30050B101 106 14,688 SH   OTR 1 10,457 4,231 0
EVOLENT HEALTH INC COMMON STOCK 30050B101 5 733 SH   DFND 5 733 0 0
EVOQUA WATER TECHNOLOGIES CO COMMON STOCK 30057T105 2 139 SH   DFND 1 139 0 0
EXACT SCIENCES CORP COMMON STOCK 30063P105 9,139 101,124 SH   DFND 1 100,024 0 1,100
EXACT SCIENCES CORP COMMON STOCK 30063P105 172 1,896 SH   OTR 1 1,431 199 266
EXANTAS CAPITAL CORP COMMON STOCK 30068N105 24 2,098 SH   DFND 1 389 0 1,709
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ROBO GLOBAL ROBOTICS ETF OTHER 301505707 40 1,035 SH   OTR 1 1,035 0 0
EXELON CORPORATION COMMON STOCK 30161N101 4,672 96,719 SH   DFND 1 93,510 0 3,209
EXELON CORPORATION COMMON STOCK 30161N101 1,796 37,175 SH   OTR 1 34,704 441 2,030
EXELON CORPORATION COMMON STOCK 30161N101 24 512 SH   DFND 5 512 0 0
EXELIXIS INC COMMON STOCK 30161Q104 227 12,851 SH   DFND 1 12,041 0 810
EXELIXIS INC COMMON STOCK 30161Q104 160 9,075 SH   OTR 1 0 0 9,075
EXLSERVICE HOLDINGS INC COMMON STOCK 302081104 1,253 18,716 SH   DFND 1 14,906 0 3,810
EXLSERVICE HOLDINGS INC COMMON STOCK 302081104 92 1,378 SH   OTR 1 1,378 0 0
EXLSERVICE HOLDINGS INC COMMON STOCK 302081104 29 445 SH   DFND 5 445 0 0
EXPEDIA INC COMMON STOCK 30212P303 1,625 12,091 SH   DFND 1 11,791 0 300
EXPEDIA INC COMMON STOCK 30212P303 589 4,386 SH   OTR 1 1,229 787 2,370
EXPEDIA INC COMMON STOCK 30212P303 42 315 SH   DFND 5 315 0 0
EXPEDITORS INTL WASH INC COMMON STOCK 302130109 9,168 123,403 SH   DFND 1 123,347 0 56
EXPEDITORS INTL WASH INC COMMON STOCK 302130109 6,529 87,890 SH   OTR 1 81,216 4,525 2,149
EXPEDITORS INTL WASH INC COMMON STOCK 302130109 13 183 SH   DFND 5 183 0 0
EXPONENT INC COMMON STOCK 30214U102 836 11,957 SH   DFND 1 8,556 0 3,401
EXPONENT INC COMMON STOCK 30214U102 417 5,959 SH   OTR 1 1,128 1,111 3,720
EXPONENT INC COMMON STOCK 30214U102 97 1,396 SH   DFND 5 1,396 0 0
EXTENDED STAY AMERICA INC COMMON STOCK 30224P200 5 315 SH   DFND 1 315 0 0
EXTRA SPACE STORAGE INC COMMON STOCK 30225T102 3,400 29,108 SH   DFND 1 28,844 0 264
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EZCORP INC CL A COMMON STOCK 302301106 9 1,435 SH   DFND 1 0 0 1,435
EXXON MOBIL CORP COMMON STOCK 30231G102 239,173 3,387,228 SH   DFND 1 2,932,906 0 454,322
EXXON MOBIL CORP COMMON STOCK 30231G102 159,539 2,259,449 SH   OTR 1 2,162,967 46,546 49,936
EXXON MOBIL CORPORATION COMMON STOCK 30231G102 581 8,237 SH   DFND 5 7,986 0 251
EYEGATE PHARMACEUTICALS INC COMMON STOCK 30233M503 9 2,710 SH   OTR 1 2,710 0 0
FBL FINL GROUP INC CL A COMMON STOCK 30239F106 16 261 SH   DFND 1 0 0 261
FLIR SYSTEMS INC COMMON STOCK 302445101 6,050 115,032 SH   DFND 1 112,649 0 2,383
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F M C CORPORATION COMMON STOCK 302491303 1,476 16,826 SH   DFND 1 16,246 0 580
F M C CORPORATION COMMON STOCK 302491303 334 3,804 SH   OTR 1 3,080 529 195
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FARO TECHNOLOGIES INC COMMON STOCK 311642102 41 838 SH   DFND 1 823 0 15
FARO TECHNOLOGIES INC COMMON STOCK 311642102 28 562 SH   OTR 1 95 467 0
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FEDERATED INVS INC COMMON STOCK 314211103 8 267 SH   OTR 1 0 118 149
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FED EX CORP COMMON STOCK 31428X106 6,198 42,578 SH   OTR 1 40,501 202 1,875
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FERRELLGAS PARTNERS LP COMMON STOCK 315293100 5 4,405 SH   DFND 1 4,405 0 0
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F5 NETWORKS INC COMMON STOCK 315616102 571 4,068 SH   OTR 1 1,924 84 2,060
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FIBROGEN INC COMMON STOCK 31572Q808 80 2,172 SH   OTR 1 0 0 2,172
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FIFTH THIRD BANCORP COMMON STOCK 316773100 13,226 483,049 SH   DFND 1 470,608 0 12,441
FIFTH THIRD BANCORP COMMON STOCK 316773100 6,923 252,866 SH   OTR 1 251,422 1,444 0
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51JOB INC A D R COMMON STOCK 316827104 182 2,454 SH   DFND 1 2,288 0 166
51JOB INC A D R COMMON STOCK 316827104 12 171 SH   OTR 1 153 0 18
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FIREEYE INC COMMON STOCK 31816Q101 821 61,541 SH   DFND 1 50,163 0 11,378
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FIRST AMERICAN FINANCIAL COMMON STOCK 31847R102 148 2,494 SH   OTR 1 2,000 47 447
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FIRST BUSINESS FINL SVCS INC COMMON STOCK 319390100 48 2,000 SH   DFND 1 2,000 0 0
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FIRST FINANCIAL BANCORP COMMON STOCK 320209109 1,819 74,328 SH   OTR 1 74,328 0 0
FIRST FINL BANKSHARES INC COMMON STOCK 32020R109 350 10,502 SH   DFND 1 1,302 0 9,200
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FIRST FINL CORP IND COMMON STOCK 320218100 65 1,500 SH   DFND 1 1,500 0 0
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FIRST HORIZON NATIONAL CORP COMMON STOCK 320517105 198 12,212 SH   DFND 1 4,208 0 8,004
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FIRST HAWAIIAN INC COMMON STOCK 32051X108 3 131 SH   DFND 5 131 0 0
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FIRST INDL RLTY TR INC COMMON STOCK 32054K103 328 8,273 SH   OTR 1 6,563 1,710 0
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FIRST MERCHANTS CORP COMMON STOCK 320817109 756 20,081 SH   DFND 1 20,081 0 0
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FIRST REPUBLIC BANK SAN FRAN COMMON STOCK 33616C100 1,537 15,897 SH   OTR 1 6,039 0 9,858
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FIRST SOLAR INC COMMON STOCK 336433107 412 7,095 SH   OTR 1 7,095 0 0
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FIRST TRUST DOW JONES SELECT OTHER 33718M105 2 56 SH   DFND 5 0 0 56
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FIRST TRUST ISE WATER IDX ETF OTHER 33733B100 9,196 163,884 SH   DFND 1 161,385 0 2,499
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FIRST TRUST CAPITAL STRENGTH OTHER 33733E104 651 11,470 SH   DFND 5 1,098 0 10,372
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FIRST TRUST AMEX BIO INDEX OTHER 33733E203 10 86 SH   DFND 5 70 0 16
FIRST TRUST DOW JONES INTERNET OTHER 33733E302 375 2,783 SH   DFND 1 2,031 0 752
FIRST TRUST DOW JONES INTERNET OTHER 33733E302 92 682 SH   OTR 1 182 0 500
FIRST TRUST DOW JONES INTERNET OTHER 33733E302 3,002 22,297 SH   DFND 5 11,519 0 10,778
FIRST TRUST NASDAQ 100 EX TECH OTHER 33733E401 49 925 SH   DFND 1 925 0 0
FIRST TRUST MULTI CAP GROWTH E OTHER 33733F101 7 100 SH   DFND 1 100 0 0
FIRST TR NASDQ 100 EQUAL WGHTD OTHER 337344105 151 2,276 SH   DFND 1 1,276 0 1,000
FIRST TR NASDQ 100 EQUAL WGHTD OTHER 337344105 100 1,500 SH   OTR 1 1,500 0 0
FIRST TRUST NASDAQ 100 EQUAL OTHER 337344105 16 251 SH   DFND 5 0 0 251
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FIRST TR VALUE LINE DVD INDX F OTHER 33734H106 404 11,586 SH   DFND 1 11,586 0 0
FIRST TR VALUE LINE DVD INDX F OTHER 33734H106 65 1,850 SH   OTR 1 1,850 0 0
FIRST TRUST VALUE LINE DVD OTHER 33734H106 17,614 505,161 SH   DFND 5 121,567 0 383,594
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FIRST TRUST LARGE CAP CORE ETF OTHER 33734K109 29 480 SH   OTR 1 480 0 0
FIRST TRUST LARGE CAP CORE OTHER 33734K109 3,368 55,305 SH   DFND 5 1,146 0 54,159
FIRST TRUST CONSUMER DISCRET E OTHER 33734X101 21 490 SH   DFND 1 490 0 0
FIRST TRUST CONSUMER STAPLES E OTHER 33734X119 44 901 SH   DFND 1 901 0 0
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FIRST TRUST ENERGY ALPHADEX ET OTHER 33734X127 10 1,063 SH   DFND 1 1,063 0 0
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FIRST TRUST TECHNOLOGY ALPHA E OTHER 33734X176 97 1,491 SH   DFND 1 1,491 0 0
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FIRST TRUST UTILITIES ALPHAD E OTHER 33734X184 15 500 SH   DFND 1 500 0 0
FIRST TRUST CLOUD COMPUTING ET OTHER 33734X192 385 6,820 SH   DFND 1 6,820 0 0
FIRST TRUST CLOUD COMPUTING ET OTHER 33734X192 240 4,250 SH   OTR 1 4,250 0 0
FIRST TRUST ISE CLOUD COMPUTNG OTHER 33734X192 45 804 SH   DFND 5 0 0 804
FIRST TRUST DOW JONES GL SEL E OTHER 33734X200 117 5,100 SH   OTR 1 5,100 0 0
FIRST TR SML CP CORE ALPHA F E OTHER 33734Y109 18 300 SH   DFND 1 300 0 0
FIRST TR SM CAP CORE ALPHADEX OTHER 33734Y109 201 3,340 SH   DFND 5 0 0 3,340
FIRST TRUST MID CAP CORE ALP E OTHER 33735B108 68 1,008 SH   DFND 1 1,008 0 0
FIRST TRUST MID CAP CORE OTHER 33735B108 24 360 SH   DFND 5 189 0 171
FIRST TRUST LARGE CAP VALUE ET OTHER 33735J101 151 2,935 SH   DFND 1 2,935 0 0
FIRST TRUST LARGE CAP VALUE OTHER 33735J101 84 1,627 SH   DFND 5 0 0 1,627
FIRST TRUST LARGE CAP GR ETF OTHER 33735K108 186 2,664 SH   DFND 1 2,664 0 0
FIRST TRUST LRG CAP GROWTH OPP OTHER 33735K108 867 12,401 SH   DFND 5 10,555 0 1,846
FIRST TRUST DOW JONES STOXX OTHER 33735T109 55 4,338 SH   DFND 5 0 0 4,338
FIRST TR GLB WIND ENGY ETF OTHER 33736G106 211 16,055 SH   DFND 1 16,055 0 0
FIRST TR GLB WIND ENGY ETF OTHER 33736G106 20 1,500 SH   OTR 1 1,500 0 0
FIRST TRUST NASDAQ ABA COMMNTY OTHER 33736Q104 2 50 SH   DFND 5 50 0 0
FIRST TRST NASD CL EDG SGIIF E OTHER 33737A108 5 100 SH   DFND 1 100 0 0
FIRST TRUST NASDAQ CLEAN EDGE OTHER 33737A108 6 113 SH   DFND 5 0 0 113
FIRST TR EUROPE ALPHADEX FUND OTHER 33737J117 328 9,561 SH   DFND 5 0 0 9,561
FT BRAZIL ALPHADEX FUND OTHER 33737J133 1 36 SH   DFND 5 0 0 36
FT SOUTH KOREA ALPHADEX FUND OTHER 33737J166 0 19 SH   DFND 5 0 0 19
FT DEVELOPED MARKETS EX-US OTHER 33737J174 48 926 SH   DFND 5 0 0 926
FT EMERGING MARKETS ALPHADEX OTHER 33737J182 1,156 49,133 SH   DFND 5 0 0 49,133
FIRST TRUST MID CAP GROWTH ETF OTHER 33737M102 71 1,589 SH   DFND 1 1,589 0 0
FT SMALL CAP GROWTH ALPHADEX OTHER 33737M300 13 298 SH   DFND 5 0 0 298
FT SMALL CAP VALUE ALPHADEX FD OTHER 33737M409 856 24,696 SH   DFND 5 0 0 24,696
FIRST TRUST NORTH AMERICAN ETF OTHER 33738D101 24 950 SH   DFND 1 950 0 0
FIRST TRUST SENIOR LOAN ETF OTHER 33738D309 16 332 SH   DFND 1 332 0 0
FIRST TRUST SENIOR LOAN OTHER 33738D309 1,401 29,547 SH   DFND 5 0 0 29,547
FIRST TRUST HIGH YIELD ETF OTHER 33738D408 13 268 SH   DFND 1 268 0 0
FIRST TRUST HIGH YIELD LONG OTHER 33738D408 2,399 49,732 SH   DFND 5 0 0 49,732
FIRST TRUST MULTI ASSET DIV ET OTHER 33738R100 33 1,800 SH   DFND 1 0 0 1,800
FT OTHER 33738R118 187 4,665 SH   DFND 5 0 0 4,665
FIRST TRUST EXCHANGE TRADED FU OTHER 33738R308 7 300 SH   DFND 1 300 0 0
FIRST TRUST HIGH INCOME OTHER 33738R308 144 6,420 SH   DFND 5 0 0 6,420
FIRST TRUST LOW BETA INCOME OTHER 33738R407 15 693 SH   DFND 5 0 0 693
FT NASDAQ RISING DVD ACHIEVERS OTHER 33738R506 3,030 95,096 SH   DFND 5 0 0 95,096
FT DORSEY WRIGHT FOCUS FIVE OTHER 33738R605 103 3,504 SH   DFND 5 1,837 0 1,667
FIRST TRUST NASDAQ SEMI- OTHER 33738R811 5 150 SH   DFND 5 0 0 150
FIRST TRUST DORSEY WRIGHT INTL OTHER 33738R886 16 836 SH   DFND 1 836 0 0
FIRST TRUST PREFERRED SEC INC OTHER 33739E108 270 13,687 SH   DFND 1 13,687 0 0
FT PREFERRED SECURITIES & INC OTHER 33739E108 964 48,877 SH   DFND 5 0 0 48,877
FIRST TRUST MANAGED MUNICIPAL OTHER 33739N108 20 368 SH   DFND 1 368 0 0
FIRST TRUST MANAGED MUNICIPAL OTHER 33739N108 85 1,530 SH   DFND 5 858 0 672
FIRST TRUST LOW DURATION MTGE OTHER 33739Q200 3,939 75,766 SH   DFND 5 0 0 75,766
FIRST TRUST ENHANCED SHORT OTHER 33739Q408 2,806 46,710 SH   DFND 5 1,168 0 45,542
FIRST TRUST TCW OPPORTUNISTIC OTHER 33740F805 1,206 22,910 SH   DFND 5 0 0 22,910
FIRSTCASH INC COMMON STOCK 33767D105 238 2,594 SH   DFND 1 2,594 0 0
FIRSTCASH INC COMMON STOCK 33767D105 84 916 SH   OTR 1 916 0 0
FIRSTSERVICE CORP COMMON STOCK 33767E202 88 857 SH   DFND 1 834 0 23
FIRSTSERVICE CORP COMMON STOCK 33767E202 134 1,310 SH   OTR 1 0 140 1,170
FISERV INC COMMON STOCK 337738108 28,224 272,450 SH   DFND 1 261,462 0 10,988
FISERV INC COMMON STOCK 337738108 6,296 60,779 SH   OTR 1 47,471 10,504 2,804
FISERV INC COMMON STOCK 337738108 87 843 SH   DFND 5 843 0 0
FLAGSTAR BANCORP INC COMMON STOCK 337930705 106 2,850 SH   DFND 1 2,850 0 0
FIRST ENERGY CORP COMMON STOCK 337932107 1,932 40,060 SH   DFND 1 38,417 0 1,643
FIRST ENERGY CORP COMMON STOCK 337932107 629 13,046 SH   OTR 1 12,911 86 49
FIRSTENERGY CORP COMMON STOCK 337932107 6 138 SH   DFND 5 138 0 0
FITBIT INC A COMMON STOCK 33812L102 40 10,410 SH   DFND 1 10,410 0 0
FIVE BELOW COMMON STOCK 33829M101 27,595 218,834 SH   DFND 1 217,969 0 865
FIVE BELOW COMMON STOCK 33829M101 3,240 25,700 SH   OTR 1 25,266 354 80
FIVE BELOW INC COMMON STOCK 33829M101 144 1,146 SH   DFND 5 1,146 0 0
FIVE9 INC COMMON STOCK 338307101 455 8,466 SH   DFND 1 8,466 0 0
FIVE9 INC COMMON STOCK 338307101 160 2,980 SH   OTR 1 1,240 990 750
FIVE POINT HOLDINGS LLC CL A COMMON STOCK 33833Q106 89 11,900 SH   DFND 1 11,900 0 0
FLAHERTY CRUMRINE PREFERRED OTHER 338478100 50 2,386 SH   DFND 1 2,386 0 0
FLAHERTY CRUMRINE PREFERRED OTHER 338478100 275 13,132 SH   OTR 1 13,132 0 0
FLAHERTY CRUMRINE PREFERRED AN OTHER 338480106 4 300 SH   DFND 1 300 0 0
FLAHERTY & CRUMRINE OTHER 33848E106 417 34,117 SH   DFND 5 17,463 0 16,654
FLEETCOR TECHNOLOGIES INC COMMON STOCK 339041105 739 2,577 SH   DFND 1 2,375 0 202
FLEETCOR TECHNOLOGIES INC COMMON STOCK 339041105 1,100 3,836 SH   OTR 1 3,462 296 78
FLEXSHARES MSTAR EMRG MKTS FCT OTHER 33939L308 11 235 SH   DFND 1 235 0 0
FLEXSHARES GLOBAL UPSTREAM N E OTHER 33939L407 302 9,685 SH   DFND 1 9,685 0 0
FLEXSHARES GLBL QUALITY REAL E OTHER 33939L787 11 165 SH   DFND 1 165 0 0
FLEXSHARES GLBL QUALITY REAL E OTHER 33939L787 11 175 SH   OTR 1 175 0 0
FLEXSHARES STOXX GLOBAL BROAD OTHER 33939L795 13 258 SH   DFND 1 258 0 0
FLEXSHARES INTL QUAL DIV ETF OTHER 33939L837 7 335 SH   DFND 1 335 0 0
FLEXSHARES QUALITY DIVIDEND ET OTHER 33939L845 17 368 SH   DFND 1 368 0 0
FLEXSHARES QUALITY DIVIDEND ET OTHER 33939L845 282 6,173 SH   OTR 1 6,173 0 0
FLEXSHARES QUALITY DIVIDEND ET OTHER 33939L860 3 68 SH   DFND 1 68 0 0
FLEXSHARES QUALITY DIVIDEND ET OTHER 33939L860 306 6,734 SH   OTR 1 6,734 0 0
FLOOR DECOR HOLDINGS INC A COMMON STOCK 339750101 432 8,438 SH   DFND 1 8,438 0 0
FLOOR DECOR HOLDINGS INC A COMMON STOCK 339750101 94 1,844 SH   OTR 1 1,392 452 0
FLUOR CORP COMMON STOCK 343412102 1,198 62,609 SH   DFND 1 62,587 0 22
FLUOR CORP COMMON STOCK 343412102 304 15,891 SH   OTR 1 15,891 0 0
FLOWERS FOODS INC COMMON STOCK 343498101 1,346 58,222 SH   DFND 1 58,222 0 0
FLOWERS FOODS INC COMMON STOCK 343498101 245 10,614 SH   OTR 1 6,070 88 4,456
FLOWSERVE CORP COMMON STOCK 34354P105 315 6,742 SH   DFND 1 6,694 0 48
FLOWSERVE CORP COMMON STOCK 34354P105 247 5,295 SH   OTR 1 5,173 122 0
FLUIDIGM CORP COMMON STOCK 34385PAA6 287 3,000 SH   DFND 5 3,000 0 0
FLUSHING FINANCIAL CORPORATION COMMON STOCK 343873105 2 84 SH   DFND 1 84 0 0
FLUSHING FINANCIAL CORPORATION COMMON STOCK 343873105 0 18 SH   OTR 1 0 18 0
FLY LEASING LTD A D R COMMON STOCK 34407D109 5,636 274,755 SH   DFND 1 273,089 0 1,666
FLY LEASING LTD A D R COMMON STOCK 34407D109 1,093 53,312 SH   OTR 1 52,934 378 0
FOMENTO ECONOMICO MEX SP A D R COMMON STOCK 344419106 3,833 41,862 SH   DFND 1 41,256 0 606
FOMENTO ECONOMICO MEX SP A D R COMMON STOCK 344419106 686 7,488 SH   OTR 1 7,168 294 26
FOMENTO ECONOMICO MEX SAB D CV COMMON STOCK 344419106 15 167 SH   DFND 5 167 0 0
FOOT LOCKER INC COMMON STOCK 344849104 15,415 357,150 SH   DFND 1 356,110 0 1,040
FOOT LOCKER INC COMMON STOCK 344849104 2,004 46,453 SH   OTR 1 46,016 52 385
FOOT LOCKER INC COMMON STOCK 344849104 2 47 SH   DFND 5 47 0 0
FORD MOTOR CO COMMON STOCK 345370860 6,054 660,923 SH   DFND 1 640,601 0 20,322
FORD MOTOR CO COMMON STOCK 345370860 971 106,000 SH   OTR 1 93,000 0 13,000
FORD MOTOR CO COMMON STOCK 345370860 7 863 SH   DFND 5 516 0 347
FORESCOUT TECHNOLOGIES INC COMMON STOCK 34553D101 782 20,614 SH   DFND 1 20,514 0 100
FORESCOUT TECHNOLOGIES INC COMMON STOCK 34553D101 228 6,007 SH   OTR 1 6,007 0 0
FORESTAR GROUP INC COMMON STOCK 346232101 5 279 SH   DFND 1 279 0 0
FORMFACTOR INC COMMON STOCK 346375108 183 9,833 SH   OTR 1 9,833 0 0
FORRESTER RESH INC COMMON STOCK 346563109 24 750 SH   DFND 1 750 0 0
FORRESTER RESH INC COMMON STOCK 346563109 9 275 SH   OTR 1 275 0 0
FORTIS INC COMMON STOCK 349553107 424 10,010 SH   DFND 1 7,267 0 2,743
FORTIS INC COMMON STOCK 349553107 122 2,879 SH   OTR 1 2,785 0 94
FORTINET INC COMMON STOCK 34959E109 1,251 16,299 SH   DFND 1 16,208 0 91
FORTINET INC COMMON STOCK 34959E109 363 4,734 SH   OTR 1 2,754 11 1,969
FORTIVE CORP WI COMMON STOCK 34959J108 6,680 97,444 SH   DFND 1 95,111 0 2,333
FORTIVE CORP WI COMMON STOCK 34959J108 1,629 23,763 SH   OTR 1 16,403 944 6,416
FORTIVE CORP COMMON STOCK 34959J108 22 330 SH   DFND 5 330 0 0
FORTUNE BRANDS HOME & SECURI COMMON STOCK 34964C106 1,458 26,663 SH   DFND 1 25,372 0 1,291
FORTUNE BRANDS HOME & SECURI COMMON STOCK 34964C106 718 13,137 SH   OTR 1 12,020 1,099 18
FORUM ENERGY TECHNOLOGIES IN COMMON STOCK 34984V100 1 672 SH   DFND 1 672 0 0
FORWARD AIR CORP COMMON STOCK 349853101 403 6,325 SH   DFND 1 6,281 0 44
FORWARD AIR CORP COMMON STOCK 349853101 48 756 SH   OTR 1 756 0 0
FORWARD AIR CORPORATION COMMON STOCK 349853101 18 293 SH   DFND 5 293 0 0
FOSSIL GROUP INC COMMON STOCK 34988V106 3 208 SH   DFND 1 208 0 0
FOSTER L B CO CL A COMMON STOCK 350060109 447 20,633 SH   DFND 1 20,633 0 0
FOSTER L B CO CL A COMMON STOCK 350060109 81 3,777 SH   OTR 1 3,623 0 154
FOUR CORNERS PROPERTY TRUST COMMON STOCK 35086T109 256 9,051 SH   DFND 1 4,783 0 4,268
FOUR CORNERS PROPERTY TRUST COMMON STOCK 35086T109 516 18,254 SH   OTR 1 5,927 12,327 0
FOX CORP CLASS A W I COMMON STOCK 35137L105 859 27,223 SH   DFND 1 26,647 0 576
FOX CORP CLASS A W I COMMON STOCK 35137L105 189 5,979 SH   OTR 1 5,921 1 57
FOX CORP COMMON STOCK 35137L105 15 486 SH   DFND 5 486 0 0
FOX CORP CLASS B COMMON STOCK 35137L204 42 1,316 SH   DFND 1 929 0 387
FOX CORP CLASS B COMMON STOCK 35137L204 49 1,571 SH   OTR 1 1,530 0 41
FOX FACTORY HOLDING CORP COMMON STOCK 35138V102 4,255 68,359 SH   DFND 1 65,918 0 2,441
FOX FACTORY HOLDING CORP COMMON STOCK 35138V102 431 6,929 SH   OTR 1 3,693 746 2,490
FRANCO NEVADA CORP COMMON STOCK 351858105 270 2,957 SH   DFND 1 2,795 0 162
FRANCO NEVADA CORP COMMON STOCK 351858105 1,881 20,637 SH   OTR 1 20,452 56 129
FRANKLIN ELEC INC COMMON STOCK 353514102 750 15,680 SH   DFND 1 15,680 0 0
FRANKLIN ELEC INC COMMON STOCK 353514102 155 3,241 SH   OTR 1 3,241 0 0
FRANKLIN ELECTRIC CO INC COMMON STOCK 353514102 14 311 SH   DFND 5 311 0 0
FRANKLIN FINANCIAL NETWORK COMMON STOCK 35352P104 3 84 SH   DFND 1 84 0 0
FRANKLIN RES INC COMMON STOCK 354613101 1,044 36,158 SH   DFND 1 35,433 0 725
FRANKLIN RES INC COMMON STOCK 354613101 163 5,655 SH   OTR 1 2,750 2,905 0
FRANKLIN RESOURCES INC COMMON STOCK 354613101 2 94 SH   DFND 5 94 0 0
FRANKLIN STR PPTYS CORP COMMON STOCK 35471R106 2 254 SH   DFND 1 254 0 0
FRANKLIN STR PPTYS CORP COMMON STOCK 35471R106 8 1,000 SH   OTR 1 1,000 0 0
FREEPORT MCMORAN INC COMMON STOCK 35671D857 425 44,457 SH   DFND 1 41,239 0 3,218
FREEPORT MCMORAN INC COMMON STOCK 35671D857 360 37,592 SH   OTR 1 36,849 0 743
FREEPORT MCMORAN COPPER & GOLD COMMON STOCK 35671D857 2 279 SH   DFND 5 279 0 0
FREIGHTCAR AMER INC COMMON STOCK 357023100 2 419 SH   DFND 1 419 0 0
FRESENIUS AKTIENGESELLSCHAFT A COMMON STOCK 358029106 1,236 36,797 SH   DFND 1 34,828 0 1,969
FRESENIUS AKTIENGESELLSCHAFT A COMMON STOCK 358029106 180 5,340 SH   OTR 1 5,169 125 46
FRESHPET INC COMMON STOCK 358039105 492 9,869 SH   DFND 1 9,869 0 0
FRESHPET INC COMMON STOCK 358039105 63 1,274 SH   OTR 1 1,274 0 0
FRONT YARD RESIDENTIAL CORP COMMON STOCK 35904G107 4 329 SH   DFND 1 329 0 0
FRONTDOOR INC COMMON STOCK 35905A109 144 2,969 SH   DFND 1 2,969 0 0
FRONTDOOR INC COMMON STOCK 35905A109 11 222 SH   OTR 1 222 0 0
FRONTIER COMMUNICATIONS CORP COMMON STOCK 35906A306 0 648 SH   DFND 1 617 0 31
FRONTIER COMMUNICATIONS CORP COMMON STOCK 35906A306 0 229 SH   OTR 1 199 1 29
FUEL TECH, INC. COMMON STOCK 359523107 240 243,476 SH   DFND 1 0 0 243,476
FULLER H B CO COMMON STOCK 359694106 1,740 37,365 SH   DFND 1 34,857 0 2,508
FULLER H B CO COMMON STOCK 359694106 2,839 60,973 SH   OTR 1 53,514 1,829 5,630
FULTON FINL CORP PA COMMON STOCK 360271100 1 53 SH   DFND 1 53 0 0
FULTON FINL CORP PA COMMON STOCK 360271100 37 2,307 SH   OTR 1 2,307 0 0
FUTUREFUEL CORPORATION COMMON STOCK 36116M106 345 28,863 SH   DFND 1 28,863 0 0
FUTUREFUEL CORPORATION COMMON STOCK 36116M106 182 15,239 SH   OTR 1 15,239 0 0
GAMCO INVS INC COMMON STOCK 361438104 137 7,000 SH   DFND 1 7,000 0 0
G A T X CORP COMMON STOCK 361448103 1,488 19,189 SH   DFND 1 18,416 0 773
G A T X CORP COMMON STOCK 361448103 268 3,466 SH   OTR 1 3,409 0 57
GATX CORP COMMON STOCK 361448103 21 275 SH   DFND 5 275 0 0
GEO GROUP INC THE COMMON STOCK 36162J106 94 5,423 SH   DFND 1 4,251 0 1,172
GCI LIBERTY INC CLASS A COMMON STOCK 36164V305 147 2,361 SH   DFND 1 2,336 0 25
GCI LIBERTY INC CLASS A COMMON STOCK 36164V305 77 1,249 SH   OTR 1 344 839 66
GCP APPLIED TECHNOLOGIES INC COMMON STOCK 36164Y101 8 440 SH   DFND 1 440 0 0
GCP APPLIED TECHNOLOGIES INC COMMON STOCK 36164Y101 1 49 SH   OTR 1 49 0 0
GDS HOLDING LTD A D R COMMON STOCK 36165L108 275 6,858 SH   DFND 1 4,188 0 2,670
GDS HOLDING LTD A D R COMMON STOCK 36165L108 1 35 SH   OTR 1 35 0 0
GW PHARMACEUTICALS A D R COMMON STOCK 36197T103 360 3,130 SH   DFND 1 3,130 0 0
GW PHARMACEUTICALS A D R COMMON STOCK 36197T103 996 8,662 SH   OTR 1 8,620 42 0
G I I I APPAREL GROUP LTD COMMON STOCK 36237H101 474 18,358 SH   DFND 1 18,147 0 211
G I I I APPAREL GROUP LTD COMMON STOCK 36237H101 75 2,913 SH   OTR 1 1,862 1,051 0
G-III APPAREL COMMON STOCK 36237H101 6 254 SH   DFND 5 254 0 0
GABELLI EQUITY TR INC OTHER 362397101 116 19,438 SH   DFND 1 19,438 0 0
GABELLI EQUITY TR INC OTHER 362397101 29 4,822 SH   OTR 1 4,822 0 0
THE GABELLI MULTIMEDIA TRUST I OTHER 36239Q109 0 35 SH   OTR 1 35 0 0
GABELLI UTIL FD OTHER 36240A101 1 138 SH   OTR 1 138 0 0
GSI TECHNOLOGY INC COMMON STOCK 36241U106 9 1,000 SH   DFND 1 1,000 0 0
GABELLI DIVID & INCOME TR OTHER 36242H104 27 1,250 SH   DFND 1 0 0 1,250
GABELLI HEALTHCARE WELLNESS OTHER 36246K103 1 87 SH   OTR 1 87 0 0
GABELLI GLOBAL SMALL MID CAP V OTHER 36249W104 1 126 SH   DFND 1 0 0 126
GALAPAGOS NV SPON A D R COMMON STOCK 36315X101 29 190 SH   DFND 1 181 0 9
GALAPAGOS NV SPON A D R COMMON STOCK 36315X101 19 125 SH   OTR 1 65 60 0
ARTHUR J GALLAGHER CO COMMON STOCK 363576109 6,039 67,425 SH   DFND 1 67,022 0 403
ARTHUR J GALLAGHER CO COMMON STOCK 363576109 22,616 252,495 SH   OTR 1 252,454 32 9
GALLAGHER (ARTHUR J.) & CO COMMON STOCK 363576109 51 574 SH   DFND 5 574 0 0
GAMCO GLOBAL GOLD NAT RES INC OTHER 36465A109 23 5,354 SH   DFND 1 5,354 0 0
GAMCO GLOBAL GOLD NAT RES INC OTHER 36465A109 128 29,429 SH   OTR 1 29,429 0 0
GAMING & LEISURE PROPE W I COMMON STOCK 36467J108 784 20,510 SH   DFND 1 17,117 0 3,393
GAMING & LEISURE PROPE W I COMMON STOCK 36467J108 60 1,566 SH   OTR 1 1,218 109 239
GAMING AND LEISURE PROPERTIES COMMON STOCK 36467J108 12 338 SH   DFND 5 338 0 0
GAMESTOP CORP CLASS A COMMON STOCK 36467W109 21 3,716 SH   DFND 1 1,722 0 1,994
GAMESTOP CORP CLASS A COMMON STOCK 36467W109 31 5,600 SH   OTR 1 5,600 0 0
GANNETT CO INC COMMON STOCK 36473H104 8 770 SH   DFND 1 770 0 0
GANNETT CO INC COMMON STOCK 36473H104 1 50 SH   OTR 1 50 0 0
THE GAP INC COMMON STOCK 364760108 687 39,592 SH   DFND 1 36,081 0 3,511
THE GAP INC COMMON STOCK 364760108<